Collins and Kempton: Lineate Leaves 
third resulted from crossing two normal 
green plants, and the remaining three 
ears were the result of self-pollinating 
lineate plants. 
DEGREE OF LINEATION VARIABLE 
Seeds from all six ears were planted 
in 1919, and all of the progenies pro- 
duced some lineate plants. The prog- 
enies, however, were not alike, and 
it was apparent that this character was 
extremely variable in expression. In 
classifying the plants an attempt was 
made to estimate the degree of linea- 
tion. A scale of ten grades was adopted, 
re) 
accord well with the expectation based 
on the assumption that lineate is a 
simple Mendelian character recessive 
to the normal form. The two green 
plants will be tested this coming 
season, and while it is within the realm 
of possibility that they are in reality 
lineate, the fact that they occurred in 
the progeny which exhibited the most 
intense expression of lineation is an 
argument against this explanation. 
RESULTS FROM RELATED PROGENIES 
In addition to the six ears secured 
from progenies which produced lineate 
TABLE IIl.—Proportion of Green to Lineate Plants, the Average Degree of Lineation and the Average 
Leaf from the Seed on Which Lineations First Appeared. 
- : 
“ | Aver. leaf on 
Designation of Female Male eee Tea | Degree of which linea- 
progeny parent parent plants plants lineation le tions 3 
| | rst appeare 
Phi24L2L1.....|Lineate.....|Self........ fw 0 28 5.37.20 10.7 
Phi24T10L2.. 2... Lyman sos WftSalbin 2G) 0 41 27a 28) | OUR 
Phi24L10L3.....| Lineate... Selifices. oon: | 2 19 5.84+.14 | DES, 
Ph124L2L2.....|Green.. .|Lineate... . .| 15 14 4.00+.49 10.2 
Ph124L5L1...... Lineate.. Lineate. . 0 35 So (hiss 8 9.5 
Phigsniowd,.:.|\Green. .y...... Green. 054. ¢ 24 3 3.00+* 9.0 
* Probable error not calculated, since only three plants are involved in this average. 
and although the classes were arbitrary 
it was found that independent observers 
were able to place the plants in es- 
sentially the same grades. The classi- 
fication of the plants secured from the 
progenies of the six ears is shown in 
Table II. 
With the exception of the two green 
plants, found in the progeny of a self- 
pollinated lineate plant, the results 
plants in 1918, there were twelve hand- 
pollinated ears from related progenies 
which produced no lineate plants in 
1918. All of these ears are directly 
descended from the ear Ph124. 
Seeds from the twelve ears were 
planted in 1919, and the progeny of 
five of these produced some lineated 
plants. The five progenies are shown 
in Table III. 
TABLE III.—Proportion of Green to Lineate Plants in the Progeny of Five Hand-pollinated Ears. 
| | 
Saale of Female parent Male parent No. of green | No. of lineate 
L2H) O00 0 ea Greene eartec cre eli ¢. facrot eas 69 3 
121122 Wa 7A 9h ee aera Greener tices: Selfpstearadn saat 18 7 
IPHIQ4ATISLI. 2. iso Green’ aan, Self 5) snchcasewdl 7 3 
BRIQAT ZOU. 4 3d nu Green Selfie cttiseia 16 9 
WAI 2CUOT A. «4 es en Green 2. .-).5 C.at¢ Greentncnkieat 124 1 
