134 SAXIFRAGER. [Donatia. 
A large and polymorphous order, very difficult to define. The herbaceous 
genera are mainly found in the temperate regions of the Northern Hemisphere, 
or on the mountains of the tropics ; the arborescent ones have their headquarters 
in South America or Australia, with a few outlying species in Africa or Asia. 
Genera about 75; species under 600. The properties of the order are unim- 
portant. Of the 6 genera found in New Zealand, Carpodetus and Ixerba are 
monotypic and endemic; Ackama and Quintinia extend to Australia; Donatia 
has one species in New Zealand and Tasmania, and another in Fuegia; while 
Weinmunna has a wide distribution in warm climates. 
* Herbs, forming compact patches. Leaves densely imbricate. Flowers 
solitary, sessile. 
Flowers white, 4in. diam. Calyx-lobes and petals 5. 
Stamens 2. Ovary inferior, 2-3-celled bs .. 1. Donati. 
** Trees. Leaves alternate, simple, exstipulate. Stamens usually as many 
as the petals. 
Flowers racemose, small. Petals imbricate. Ovary in- 
ferior ee a 3 fe ee ... 2. QUINTINIA. 
Flowers panicled, large. Petals imbricate. Ovary superior 3. IXERBA. 
Flowers panicled, small. Petals valvate. Ovary inferior 4. CARPODETUS. 
*** Trees. Leaves opposite, stipulate. Stamens usually twice as many as 
the petals. 
Flowers panicled. Calyx valvate Sc Se -. 5. ACKAMA. 
Flowers racemose. Calyx imbricate .. ne .. 6. WEINMANNIA. 
1. DONATIA, Forst. 
Small densely tufted herbs, forming hard compact masses. 
Leaves densely imbricated, linear, coriaceous, quite entire. Flowers 
terminal, solitary, sessile, white. Calyx-tube adnate to the ovary, 
obconic ; lobes 5-7, equal or unequal. Petals 5-10, linear or ovate. 
Stamens 2 or 3, inserted on the middle of an epigynous disc, and 
adnate to the base of the styles; filaments subulate or filiform ; 
anthers didymous, extrorse. Ovary inferior, 2- or 3-celled; styles 
2 or 3, short and thick or subulate, recurved; stigmas simple or 
capitellate ; ovules numerous, affixed to placentas which are pendu- 
lous from the inner angle of the cells. Capsule turbinate, indehis- 
cent, 2- or 3-celled. Seeds few in each cell, pendulous, obliquely 
ovoid ; testa membranous ; albumen fleshy; embryo small, remote 
from the hilum. 
A genus of two species, one found in New Zealand and Tasmania, the other 
a native of Fuegia. Its exact systematic position is very doubtful; it was re- 
ferred to Sawifragee by Hooker, who, however, also pointed out its affinity 
with the Stylidie@, with which it agrees in the stamens being placed on the 
centre of an epigynous disc, in the extrorse anthers, and in the placentation. 
It was removed to that order by the late Baron Mueller (‘‘ Nuovo Giornale 
Botanico Italiano,’’ xi., July, 1879). On the other hand, both Baillon and 
Engler retain it among the Saxifrages, the latter (‘‘ Naturlichen Pflanzenfami- 
lien,” Teil iii. Abt. ii.a, p. 67) constituting it a new subsection of the order. 
1. D. nove-zealandiz, Hook. f. Fl. Nov. Zel. i. 81, t. 20.— 
Stems short, 1—3in. high, densely tufted, forming broad compact 
masses in mountain-bogs. Leaves very numerous, imbricated in 
many series and clothing the entire stem and branches, erect, 
