FLOWER AND SEED COLOUR IN LUPINUS 303 

There is one important point of difference, however, between the 
flowers of the whites and those of the tinged types, viz. the colora- 
tion of the latter ones even in early stages. The young flowers of the 
whites show no trace of colour; the sepals, bracts and axis are 
quite green. 
THE SEED COLOUR. 
The seed colour of Lupinus angustifolius depends on the presence 
of pigment in the palisades of the seed coat, according to the investi- 
gations of Kasanus and FruwirtH. The figure reproduced in the 
work of Fruwirtu (1915, pag. 218) shows that the pigment layer is de- 
fined to the centre of the palisade cells. The ordinary colour of the 

a (7/1) b (?/1) 
Marbled; white flecks, dark areas Non-marbled; white flecks, dark 
composed of closely packed pigment areas as in a, the lighter areas 
grains, the lighter areas with scatt- lacking. 
ered grains. 
Biel: 
seed is this (Pl. II, fig. 8). On a lighter or darker coloured pearl-gray 
ground there is found an earth-brown marbling of varying limits 
together with white flecks sharply defined. When examined under a 
magnifying lens the characteristics of this coloration will be found 
(see fig. 1). The pigment grains are collected in small groups; the 
heaping up of these groups or the scattered occurrence of the groups 
give rise to the above mentioned characteristic colour schemes as 
shown in the figure. They are very closely heaped up in the dark 
marbled areas, they are more scattered in the pearl-gray fields, and 
in the white parts only sporadic dots are found. The limits between 
the different coloured areas are remarkably sharp. I have characteri- 
zed this type as the earth-brown, marbled. Kasanus mentions three 
