I 



CARBOITIFEROUS LYCOPODS AND SPHEKOPHYLLS. 105 



1884. Sigillariostrobns, Zeiller. Ann. d. Science Nat., 6° Ser. Bot., 



Vol. XIX., p. 256. 

 1888. Sigillariostrobus, Zeiller. Flore foss. Bassin houil. d. 



Valenciennes, p. 391. 

 1897. Sigillariostrobiis, Kidston. Trans. Roy. Sac. Edin., Vol. 



XXXIX, Part i., p. 49. 



Cones sessile or borne on deciduous peduncles with few leaves, 

 or the leaves reduced to bract-like structures, cylindrical, composed 

 of a ligneous axis bearing spirally arranged sporophylls. Bracts 

 entire or ciliate at margin, rhomboidal acute, or lanceolate, 

 with an expanded rhomboidal base, caducous. Sporangia con- 

 tained in the hollow inflated base of the sporophyll. Macrospores 

 0-75 mm. to 2-00 mm. in diameter, surface smooth or covered 

 with small apiculi. Under surface marked with three ridges 

 which radiate from the centre. Microsporangia probably present. 



Remarks. — Goldenberg was the first to refer to Sigillaria certain 

 cones as their fructification.' That figured on his PI. X., fig. 1, he 

 thought might be referable to Sigillaria tessellata, and that at fig. 2 

 of the same plate to Sigillaria intermedia. Some other figures of 

 portions of cones are also given by the same author.- For these 

 fructifications Schimper founded the genus Sigillariostrobus. 



The reference of these cones to Sigillaria was suggested 

 chiefly on the e\'idence of their association with stems of that 

 genus — evidence of a very inconclusive nature ; and it was 

 not till the discovery by Mons. Zeiller thirty years later of a 

 similar cone attached to a pedicel bearing Sigillarian leaf-scars 

 that Goldenberg's cones could be accepted as undoubtedly the 

 fructification of Sigillaria.^ 



This discovery not only proved the accuracy of Goldenberg's 

 conclusions as to the position he assigned to his fossils, but also 

 proved that Sigillaria is Lycopodiaceous, an opinion by no means 

 universally accepted. Goldenburg further stated his belief that 



1 Goldenberg, Flora sarwp. foss. Heft. I. (1855); Heft. II. (1857). 



2 Goldenberg, ;. c, PI. B., figs. 18-25; PI. IV., fig. 3; Heft. II., p. 1. 



* Zeiller, Sur des cOnes de fructification dt Slgillairfs. Comptes rendus, 

 30th June, 18S4. C6nes de fructification de Sigillaires. Ann. d. Scienc. 

 Nat., 6= Ser. Bot., Vol. XIX., pp. 256-280, Pis. XI-XII., 1884. Flore foss. 

 Bassin houil. d. Vahnciennes, pp. 591-608, Pis. LXXXIX-XC, 1886-88. 



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