Little and Jones: Inheritance of Color in Great Danes 315 



Crossing E animals inter se. 



(b) Expected on 3 : 1 ratio 13 7-46 ±3. 9 



(c) Expected on 2 : 1 ratio 122-61 ±4.3 

 Difference (a) and (b) 3 ±5. 4 

 Difference (a) and (c) 18 ±5. 7 



Table IV shows the results of mating 

 •dilute animals inter se. While the 

 numbers are vSmall the results are con- 

 sistent with the supposition that dilute 

 forms are hypostatic to intense and all 

 fifty-three animals obtained are dilute 

 pigmented. 



Table IV 

 Matings D and d 



Ratio of E types with- 

 in a mixed population 



Ratio of E to Eitoe' 



animals obtained by 



random matings 



Ratio of E tvpes with- ^^*^° °^ ^^^-^'i"; ^' 

 in a mixed population ^"^^^j^ obtained by 

 ^ ^ random matmgs 



Table V shows a combination of all 

 matings involving the factor E. If 

 the E X E matings be first considered, 

 it will be noted that the ratio of 309E 

 to 22E^ to 2e^ approximates very closely 

 the numbers expected on a 75 to 8 to 1 

 distribution. Referring to the list of 

 matings given above, it will be seen 

 that the 75 : 8 : 1 ratio depends upon 

 the supposition that EE and EE^ 

 individuals occur with equal frequency 

 in a mixed population and which are 

 each of them twice as numerous as 

 Ee^ animals. This is not at all un- 

 likely, for blacks or harlequins (E) 

 and brindles (E^) are about equally 

 popular in Great Dane breeding, and 

 both of them appear far more frequently 

 than do fawns (e^). 



Table V 



Matings of EEi and ei 



Cross EXE 



{h) The factors E for full extension, E^ 



for partial extension, and e^ for 



restriction of black pigment 



In the mixed population of Great 



Danes under consideration, the ratio 



of E, El, and ei types obtained when E 



forms are crossed inter se or with ei 



will be characteristically distinct and 



will give some idea as to the degree of 



inbreeding and selection involved. Thus : 



