Little and Jones: Inheritance of Color in Great Danes 317 



that the pecuHar striping disappears, 

 and in that case the introduction of a 

 good fawn into the strain is advisable." 

 It seems therefore entirely probable 

 that at least a part of the nine blacks 

 (E) appearing in the E^xE^ matings 

 and the one black appearing in the 

 E^xei matings represent brindles of 

 this extremely dark or blackened type, 

 while the four brindles (E^ which 

 appear in matings of fawn x fawn may 

 well be due to the fact that one of their 

 parents was an extremely light brindle 

 in which the black was so reduced that 

 it appeared phenotypically like a fawn 

 and was so recorded. The occurrence 

 of exceptions of this type is therefore 

 expected, provided that the brindle 

 pattern of Great Danes behaves as all 

 other similar pattern factors hitherto 

 studied in mammals. 



(c) Factor H for harlequin spotting, 

 h for absence of harlequin spotting 

 or self coat. 



Inheritance of any form of spotting 



is complicated because of great varia- 

 bility in the degree of spot ring, and of 

 incompleteness of dominance in many 

 cases, also because genetically different 

 types of spotting all show themselves 

 as simple contrasts between colored 

 and white areas. Even with all these 

 handicaps, however, the data obtained 

 from the A. K. C. stud books indicate 

 that there are at least two genetically 

 different types of spotting in Great 

 Danes. We may first consider the more 

 striking of these, namely harlequin 

 spottings. (Plate 1, Fig. 2.) 



Animals of this type when crossed 

 together have given, in addition to 

 harlequin, a considerable number of self 

 animals as shown in Table VI. From a 

 fancier's point of view, harlequins by 

 inbreeding continually tend to grow too 

 light, that is, to have too little black 

 pigment. The method followed to cor- 

 rect this is to cross harlequin with self 

 intense blacks. (Fig. 12, number 3.) 

 This leads to an increase in black 

 areas on the resulting harlequins. 



Table VI 



Matings HxH 



Mating 



H 



A 



Totals 



Expected 3 : 1 ratio . 

 Expected 8 : 1 ratio . 

 Expected 15 : 1 ratio. 



Mating 



Matings HXh 



H 



G. . . : 



K 



Q 



V 



Bi 



Di 



Totals (a) 



Expected (aj 1 : 1 ratio. 

 Expected (b) 2 : 1 ratio. 

 Expected (d) 3 : 1 ratio. 

 Difference (a)X(b) 



Difference (a) and (c) . 

 Difference (a) and (d) 



±4.63 

 ±4.63 



±3.57 

 ±2.61 



±2.13 

 ±2.26 

 ±2.13 

 ±1.94 



Diff. 



E. Diff: 



Diff. 



= 2.2 



Matings hXh 



Mating 



P. E. of Diff, 



Diff. 

 P. E. of Diff. 



= 5.0 



= 6.5 



Totals. 



H 



515 

 43 

 44 



5 



16 

 37 

 21 

 433 

 29 



4 

 13 



8 

 13 

 24 



5 



4 



1214 



