390 



The Journal of Heredity 



now EXPECTATION \^AS FULFILLED 



Table II shows the results of crossing various types of Cocker Spaniels, made up from the 

 records of matings furnished to the authors, and based on Table I. Only those matings 

 are here shown in which the formula (class) of each parent is known from two or more 

 matings, B = black, R = red, L = Hver, Y = lemon or yellow. Figures in parenthesis indicate 

 the number litters. 



of class 3 or 4, three times to class 7, 

 and once to class 8. The results of the 

 first five matings gave thirty-six blacks 

 and twelve livers, an exact agreement 

 with the table. The next three matings, 

 (classes 3 by 7) yielded fifteen black 

 pups. The last mentioned cross (3 by 8) 

 gave four blacks and one liver. In this 

 list each of the red bitches of class 7 

 was mated more than once and in each 

 mating behaved as a class 7 should, 

 no di.seordant results being obtained. 



We have now double-checked each 

 individual which occurs in two or more 

 matings, and find that the records show 

 the actual existence of each of the 

 nine classes represented, and that with 

 two exceptions the results accord with 

 those expected from Table I. Table II 

 shows part of our actual records. 



A comparison of this table with 

 Table I will show that the results agree 

 very closely with those exjK'Cted. The 

 litter shown as a result of the mating 



of class 4 by class 4 is placed here 

 because the sire when mated to a red of 

 class 8 gave four blacks and one liver. 

 The only possible arrangement of these 

 litters is to class one as 4 by 4 and the 

 other as 4 by 8. This is, perhaps, the 

 most questionable case given in the 

 table. We have no records of the 

 matings of liver to liver. But Rev. T. 

 Moore-Smith, an extensive breeder of 

 Irish Water Sijaniels, writes us that 

 they always breed true in his kennel, and 

 so their formula is ]jrobably bE bE, 

 and they re])resent the mating 5 by vS. 



The litter shown as a resuh of two 

 livers (6 by 6) is a litter of English 

 Water Spaniels. However, there is 

 probably very little objection to its 

 inclusion since at one time both English 

 and Irish Water Sjjaniels were freel\- 

 bred with the Cocker S])aniels. 



Out of eighty-nine litters there were 

 two which could not be fitted into the 

 table. A ri'd dog of class S was su])- 



