Simpson: Coat-Pattern in Mammals 



333 



SOME COMPLICATIONS IN MENDELISM 



Emma D. 71, a Belted Red sow, sired by a pure Red Tamworth boar out of a Belted Red sow, 

 tracing back in the maternal line to a Hampshire boar on a light-roan-belted hybrid sow. 

 Although she appears true to type she is of decidedly mixed descent. The question of 

 interest to the breeder is to know how she will behave inbreeding. She was mated to a 

 Belted Red boar which was also a Tamworth-Belt hybrid, and farrowed 18 pigs. Eleven 

 were belted, while seven were recessive reds. Had the parents been of pure blood instead 

 of mongrels, all the pigs would have been expected belted. It is interesting to note that 

 the black of Hampshire has been completely eliminated, both from body and germ cells, 

 in this sow, as far as the analytical test of breeding shows. (Fig. 3.) 



scattering pigmented hairs in the selfed 

 area (the Hampshire selfing factor is 

 recessive to pure White Yorkshire) . 



The next two pigs with their standard 

 belts, pale diluted red at each end with 

 a black spot in this on top of the rump, 

 appear to have resulted from the pairing 

 of enough Tamworth-red entities from 

 the dam, which uncrossed would produce 

 a highly potent yellowish-red, which is 

 often found among the recessives of the 

 F2 Tam-York cross. The added dom- 

 inance of these two pigs was perhaps 

 received from a single Yorkshire factor, 

 with the dominant and recessive factors 

 of the Hampshire sire. 



The next two pigs, good Hampshires 

 save that their belts narrowed to zero 

 on top of the shoulders, should be classed 

 with the first three of Tam-Hamp com- 



position, only in these the selfing factor 

 was more potent against the belting. 

 The next two pigs, red with a narrow 

 belt not continuous at the withers, are 

 like the belts with yellow-red, except that 

 the hyperpot: if-, red from Tamworth, 

 by Yorkshire acceleration, is yet more 

 dominant against the Hampshire black. 

 The next one, which was nearly all red 

 with only the left fore-leg white, might 

 be classed with the last foregoing, 

 except that his belt was more incipient. 



The last three pigs, with coats all 

 white and skins mosaic, should be the 

 result of crossing nearly pure white 

 Yorkshire with the pure Hampshire. 

 Evidently none of the Tamworth selfing 

 factor was left in the conglomerate eggs. 



We learn from these experiments 

 and from direct crossing of the White 



