Kuiper: Color Inheritance in Cattle 



105 



SOME MORE PIEBALD HEIFERS USED IN THE CROSSBREEDING EXPERIMENTS 



There were more than 60 piebald heifers used in the crossing experiments with the belted bull 

 pictured in the herd on the opposite page. Six of the heifers were red and white, but the rest were 

 black and white. The coat color inheritance is illustrated by the types shown in Figure 2. (Fig. 5.) 



black around the eyes, a so-called 

 Groninger "Zwartblaard" cow. Fig. 2 

 shows some specimens together with 

 their calves. Of 54 of these animals the 

 outcome of calving could be stated: 55 

 calves were born; of these 27 bore the 

 belt character, 24 or 25 were self- 

 colored, and 3 or 4 pied. Among the 

 belted ones I found only a few with 

 the true pattern like the sire. Nearly 

 all of them had at any rate two white 

 hind feet. (Figs. 6 and 7.) In some 

 cases they showed worse deviations. 

 Of these some will be seen in Fig. 8 A-C. 

 The self-colored ones were, as a rule, 

 coal-black, but there were some among 

 them that showed a small white spot 

 on the belly, the tail-end or on the fore- 

 head. Still these animals were quite 

 distinct from even the darkest speci- 

 mens of pied cattle. Only in one case, 

 which I have not been able to examine 



personally, was it doubtful whether 

 the animal was to be set down as self- 

 colored or pied; hence the figures as 

 cited above. They were all of them 

 black-haired calves (instead of red), 

 proving that the bull was homozygous 

 with respect to black hair. The 

 "blaarkop" cow (white face with black 

 around the eyes) produced a belted 

 calf with a head like its mother. (See 

 Fig. 2.) 



Also the cross — piebald bull x belted 

 cows — has been carried out in Holland 

 of late years by Mr. Jochems at 

 Wassenaar. There, too, the calves 

 showed the three types, the propor- 

 tion, however, being entirely different, 

 viz., 18 belted, 2 self-colored, and 1 

 piebald. But we shall see presently 

 that the various cows used in these 

 experiments were probably partly 

 homozygous and partly heterozygous 



