POLLEN GRAINS OF C. COCCINEA 
Two complete grains of pollen are heré shown, enormously magnified. The dark patches in 
their centers are the nuclei, which are supposed to be the carriers of most, if not all, of the 
heredity; surrounding the nuclei is nutritive material. 
It will be noticed that the outer 
envelope of the grain is marked by thin spots, through which a pollen tube can easily break 
and grow down into the ovary of the plant to be fertilized. 
Missouri 
1. * C. Kellogit, St. Louis, Mo. 
2. * C. lanuginosa, Webb City, southwestern 
Missouri. 
3. * C. pyroformis, southeastern Missouri. 
Sargent: Middle Belt (Forms Not Listed in 
Gray) 
1. * C. Eamesii, Connecticut, rich, moist soil. 
2. * C. Neo-Londonensis, Connecticut, bor- 
ders of woods. 
3. * C. Hill, northeastern Illinois, open 
woods, river banks, rich, moist soil. 
4. * C. assurgens, northeastern Illinois, open 
woods, river banks, rich, moist soil. 
5. * C. acclivts, northeastern Illinois, banks 
of rivers at Rochester and Niagara. 
6. * C. delecta, northeastern Illinois, open 
pastures and wood borders. 
7. * C. sertata, northeastern Illinois, open 
woods, rich, moist soil. 
Note: Forms marked (*) are of purely local 
distribution. 
The forms of the group as a whole 
are seen to be either of extremely wide 
range of distribution or they are ex- 
tremely local; and these local forms 
never occur in an area not already 
covered by two or more of their more 
cosmopolitan cousins. These facts I 
consider significant. It is also of inter- 
est to note that all the new species 
recorded in Sargent but not in Gray 
and belonging to this group are confined 
to purely local distribution—two in 
(Fig. 13.) 
Connecticut, and five in northeastern 
Illinois. I venture to guess that if the 
pollen of these forms were to be investi- 
gated, it would show a high percentage 
of sterility. 
Examination of the pollen conditions 
of the Intricatae, on the other hand, 
proved them to be surprisingly sterile. 
Gray ascribes to the group six species 
and one variety. Of these seven forms 
I have investigated five: 
C. Boyntont. 
C. foetida. 
‘C. coccinea L. 
C. apposita. 
C. apposita, var. Bissell. 
Sargent describes three more forms 
not mentioned in Gray, and of these I 
was able to ascertain the pollen condi- 
tions of two: 
C. Buckleyt. 
Thus of the ten different forms that 
make up the Intricatae, I have looked 
into the spore conditions of seven and in 
no case have I found a flower that is 
not strikingly sterile. I was able to 
examine the pollen of one of the two 
extremely local species of the group— 
C. venusta, which is confined to the open 
oak and hickory woods on the dry 
273 
C. venusta. 
