120 



A. AKERMAN 



spike ^. As the generative cells arise from the tissue next under the epi- 

 dermis (Koernicke1897, p. 152) gametes with speltoid characters are not 



formed. Conse- 

 quently the off- 

 spring becomes 

 constant and of 

 the normal type. 

 Another such 

 chimaera was 

 found in a Fg-plot 

 (1917—161) from 

 a cross between 



Brown Schlan- 

 stedt spring-wheat 

 and 0715, the lat- 

 ter being a des- 

 cendant of the 

 Norwegian Bör- 

 sum spring-wheat. 



Both Brown 

 Schlanstedt and 

 Börsum are typi- 

 cal vulgare -types. 

 The plant in ques- 

 tion had five 

 straws four of 

 which had spikes 

 of the normal ty- 

 pe. The fifth spike 

 had normal outer 

 glumes on one side, 

 distinctly disten- 

 ded, while they 

 on the other side were of the speltoid type and firmly pressed to the 

 flowering scales (fig. 3). In the spring 1918 seeds were sown both from 

 the flowers next under the spelto^id outer glumes (14 seeds) and from 



Fig. 3. A cliimacra from the cross Brown Schlanstedt 

 spring-wheat X Börsumwhcat (1917 — 161). 

 (I and c. Tlie spilic with both typical and speltoid 

 outer glumes. 



b. One of the normal spikes from the same plant. 



* In order to gel conclusive evidence on this point I have undertaken a micro- 

 scopical investigation of the outeF glumes in this abnormal form and in other 

 speltoids, but hitherto no decisive results have been obtained. 



