SPELTLIKE BUD-SPORTS IN COMMON WHEAT 



125 



cally corresponded with the heterozygote, 10 seeds were sown, and of 

 these 6 plants were obtained, 5 of which were speltoids. Only one 

 belonged to the normal type. Seeds from two of the speltoids were 

 sown in the autumn 1918, and the plants originating showed also ségré- 

 gation in speltoid heterozygotes, identical with the parent plants, and 

 plants of the normal type in the ratio 1:1, as seen in table 2. 



TABLE 2. 



From these investigations of the progeny of the last described chi- 

 maera-like plant is seen, that the seeds nearest the typical outer 

 glumes in the 8 lowest spikelets gave only typical descendants, while 

 the seeds nearest the speltoid outer glumes gave rise to both speltoid- 

 heterozygotes and typical plants in the ratio 5:1. The same result 

 was obtained from seeds situated in the speltoid top of the spike. Con- 

 sequently it lies close at hand to suppose that the plant in question 

 is a real chimaera largely made up of a speltoid heterozygote together 

 with two sectors of the normal type, building up not only the epidermis 

 but also at least one cell-layer of the tissue lying next under the epi- 

 dermis \ 



This chimaera had only one straw when found in the sorting looni, 

 but fragments of another straw were seen very distinctly at the base. 

 After some searching I found a loose straw which corresponded very 

 well with the fragments. The spike of this straw had speltoid outer 

 glumes. All its seeds were sown in the autumn 1917. On account 

 of unfavourable weather conditions only 6 plants were obtained, 4 

 of which were of the normal type and 2 speltoid heterozygotes. Both 

 the typical plants and the speltoid heterozygotes resembled the ones 

 obtained from the progeny of the chimaera-like spike. 



* The segregation 1 :5 (or counted pro 2^0,33: I.«?) obliiined holli in llie 

 progeny of the lower speltoid parts of the spike and in tlie progeny of its speltoid 

 top is by no means satisfactory, as a segregation in the ratio 1:1 is to be expected. 

 Howeyer, the difference between the theoretical relation and the calculated one is not 

 even twice the standard error on account of the small number of individuals. 



