UBER EINEN FALL VON CRISS-CROSS-VERERBUNG BEI EINER ARTKREUZUNG 145 
resemble anachoreta for they receive from the ¢-bastard one curt.-X- 
chromosome, X,, as well as one anach.-X-chromosome, X,. Thus the 
dominance of the latter one gives the anachoreta-resemblance. The 
males are triploid and presumably contain also three X-chromosomes. 
The relation between allo- and autosomes in these individuals are 
then the same as in the normal diploid type. The females are also 
triploid, but these contain the allosomes XXY which makes the rela- 
tion between allo- and autosomes different from the one typical of the 
diploid female. Intersexual individuals are also found among these 
although poorly developed as a rule. The conclusion lies close at 
hand that the Y-chromosome contains certain factors which influence 
the differentiation of sex and develop specific female characteristics 
in spite of the presence of two X-chromosomes which ought to 
d-mark the individuals. The Y-chromosome naturally does not con- 
tain any larval pigment genes. No criss-cross inheritance would be 
possible if it did contain such genes. 
A Q-larva resembling anachoreta was once found in a large pro- 
geny from the cross anach.® X curt. 4. The moth of this larva was 
crossed with a male from the same hatch but no eggs were developed. 
The strange female must have arisen through non-disjunction in an egg 
of the heterogametic mother. It must have received the Y-chromo- 
some from the mother; its development into a female would otherwise 
have been impossible. The X,-chromosome must also have been ob- 
tained from the anach.-mother; the anach.-resemblance would not 
otherwise have been attained. 
\ 
ZITIERTE LITERATUR, 
1. Bateson, W. 1908. The methods and scope of genetics, an inaugural lecture 
delivered 28 Oct. 1908. Cambridge. 
2. Baur, Erwin. 1911. Einführung in die experimentelle Vererbungslehre. 1. 
Aufl. Berlin. 
3. BRIDGES, CALVIN B. 1916. Non-disjunction as proof of the chromosome theory 
of heredity. Genetics I, p. 1—52, 107—163. 
4. — 1922. The origin of variations in sexual and sex-limited characters. Amer. 
Naturalist 56, p. 51—63. 
5. DONCASTER, L. 1914. On the relations between chromosomes, sex-limited 
transmission and sex-determination in Abraxas grossulariata. Journal of 
Genetics 4, p. 1—21, pl. 1—2. 
6. Doncaster, L. and Raynor, G. H. 1906. Breeding experiments with Lepidop- 
tera. Proc. Zoolog. Soc. London. 
