STUDIES ON MENDELIAN FACTORS IN AQUILEGIA VULGARIS 189 
This analysis is by no means complete. Besides the above 
mentioned colour-segregations it would have been interesting to make 
a more complete investigation of the pleiotropism of the colour 
factors, especially of the red factors. It is possible that the pleiotropical 
effect is to be found in other parts of the plant. In any case it would 
have been interesting to investigate a greater number of plants, but 
as the chance of continuing these experiments is rather small I 
have thought it best to publish the results already at this stage. 
The flower colour in Aquilegia corresponds to the following num- 
bers of colour in KLINGSIECK et VALETTE : Code de Coleurs, viz. dark- 
blue = No. 428—429, light-blue = No. 0466—466 and red = No. 571. 
SUMMARY. 
The following factors for flower colour have been found: 
B gives light-blue flower colour. 
R gives red flower colour. 
If B and R are both present the flower becomes dark-blue; if 
they are both absent the flower becomes white. 
C develops colour all over the corolla; when absent the petals 
are white-marginated. 
The red factor R shows the following pleiotroipcal effect: 
a) The leaves become anthocyan coloured on both sides; when 
the factor R is absent the leaves have anthocyan only on the underside. 
b) The stem becomes taller. 
c) The seed-coat becomes shiny, and the endosperm turns 
darker. When the factor is absent the seed-coat is dull and dead-black. 
d) The 1000-grain weight becomes higher. 
The other factors for flower colour do not seem to have any 
pleiotropical effect. 
LITERATURE CITED. 
1. Baur, E. 1911. Untersuchungen über die Vererbung von Chromatophoren- 
merkmalen bei Melandrium, Antirrhinum und Aquilegia. Zschr. ind. Abst. 
2. — 1919. Einführung in die experimentelle Vererbungslehre. Berlin. 
Hereditas III. 13 
