E. W. BERGER ON THE CUBOMEDUSA. 65 
of only one kind of cells in the epithelium. In a tangential section 
taken just through the tips of the epithelial cells (Fig. 25) I find 
polygonal areas with a central dot. This section does not at all agree 
with Schewiakoff’s Fig. 8, in which he figures two kinds of cells. In 
Fig. 25 there can be no evidence of two kinds of cells, unless both 
kinds have like flagella, for these dots are the transverse sections of 
flagella continued within the cells (Fig. 26). 
The epithelium, then, is flagellate, a flagellum to a cell. Whether 
there are flagella on the epithelium covering the region of the con- 
cretion, I could not determine. But I believe that in all other parts, 
excepting, of course, the corneas, it is flagellated. The fibers (flagella) 
of the simple eyes are evidently the flagella of the invaginated 
epithelium. Each flagellum has a basal body, and I could in many 
instances determine that it was dumbbell-shaped (Fig. 12). This fact 
was not always evident, however, and it was only occasionally that I 
felt sure of it. Often the flagella showed only a general thickening 
within the cells (Fig. 26) while, again, the thickening (basal body) 
might be quite localized near the surface of the cell. Each flagellum 
extends into its cell, and occasionally I could trace one clear past the 
nucleus into the subepithelial nerve-tissue (Fig. 26), just as I did for 
the axial fibers of the retinal cells of the simple eyes. In those 
instances in which I could do this, the fibers could so clearly be 
traced that little if any doubt can exist. I have thus made bold 
and have drawn the flagella as continued through their cells into the 
subepithelial nerve-tissue for all the cells of the epithelium of Fig. 12. 
A word on the epithelium covering the network cells of Fig. 13. 
Conant and Schewiakoff here describe fibers from the supporting 
lamelle that pass in bundles in among the network cells. These 
fibers are supposed to be a part of the supporting lamella which 
reaches out to be a support for the epithelial cells. (Schewiakoff also 
describes similar fibers for other parts of the epithelium.) Now, as 
Conant himself shows in Fig. 13, these coarse fibers are not of the 
same consistency and staining capacity as the supporting lamella. I 
found them to stain just like the intracellular parts of the flagella or 
like the central continuations of the axial fibers of the cells of the 
simple eyes. I could, also, occasionally trace them to the surface of 
the epithelium, and beyond, when they became continued as short 
blunt processes or flagella (Fig. 13). I, therefore, conclude that they 
are sensory fibers like those I have described for the other epithelial 
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