72 JOHNS HOPKINS UNIVERSITY MORPHOLOGICAL MONOGRAPHS. 
cells, and that the smaller ones are also found in the males. This 
latter fact agrees with what I have suggested, that some of the 
floating cells arise in the ampulla. My chief reasons for my suppo- 
sition, however, are the following: I find globules of the secretion 
of the ampulla cells in some of the floating cells and also scattered 
loosely among them (Fig. 19). These globules in and among the 
floating cells have the same general appearance and a similar 
staining capacity as the secretion in the ampulla cells. Again, in 
spaces within some of the ampulla cells I find bodies resembling the 
floating cells with lumps of the secretion within them (Fig. 18). 
The conclusion, therefore, lies near that some of the floating cells 
originate within the cells of the ampulla, engulf within them some 
of the secretion, and are then expelled into the lumen of the ampulla. 
Better said, perhaps, they represent portions of the ampulla cells 
with some of the secretion. I also found several instances in which 
a floating cell had the appearance of being expelled from an ampulla 
cell. Conant suggests for a similar observation that the cells were 
about to be swallowed by the ampulla cells. I believe, however, that 
my finding a secretion similar to that within the cells of the 
ampulla, in some of the floating cells, as also bodies very much 
like them and filled with secretion within the ampulla cells, 
together with Conant’s finding floating cells in males, and finally 
the observation that the floating cells are usually quite dilapidated, 
never showing a healthy cell structure—all this leads me to conclude 
that some of the floating cells originate from the ampulla cells, and 
that they have a nutrient function in distributing the secretion. 
This is quite the reverse of what Conant supposed,—that they were 
taken in as nourishment by the ampulla cells. I also find what 
appears to be a secretion in the endoderm of the tentacles of both 
Charybdea and Tripedalia, and believe this is another source of the 
floating cells. (See below, under tentacles.) 
I also found other very darkly staining bodies (Fig. 19) both 
within the floating cells and free in the ampulla cavity, and more 
numerous in the ampulla cells themselves. This again goes to show 
that floating cells take their origin from the ampulla cells. What 
these darkly staining bodies are, I cannot say. Perhaps they are 
something akin to the “Chromatoider Nebenkorper” described by 
Lenhossek (L), or they represent another kind of secretion. If these 
floating cells are derived from the cells of the ampulla, the active 
