4 
172 BeNeEpict: GENERA OF THE FERN TRIBE VITTARIEAE 
the lines are in slight grooves or even superficial. In the P. 
lineatum type the lines are much elongated, and, with rare excep- 
tions, unconnected (PLATE 7, FIG. 8). In the pair of species which 
show this type, the sporangia are also developed in deep, partly 
covered grooves. In P. lanceolatum, a species which belongs next 
to P. lineatum, the sporangia are nearly superficial and in short, 
often more or less branching lines along the longitudinal veinlets 
(PLATE 7, FIG. 4). The specimens of P. lanceolatum figured are of 
the variety Feei. Typical P. lanceolatum is much longer and nar- 
rower in proportion, and shows soriation not so unlike P. lineatum, 
but as P. quadriseriatum and P. lineatum are situated near the 
border line between Vittaria and Polytaenium, it is not surprising 
that they should show a connecting type of soriation. In true 
Old World Antrophyum, all grades of soriation between a vittarioid 
type and the dichotomous type of A. reticulatum and A. planta- 
gineum (PLATE 7, FIG. 11), and further a practically complete 
reticulation, as in A. Boryanum, may be found. 
Polytaenium may be considered as a well-marked genus, not- 
withstanding its obvious origin from a form like Vittaria. The 
whole tribe shows evidence of being of recent origin, and if so, 
it is not. strange that the connecting links between the genera 
have been maintained. 
5. ANANTHACORUS Underwood & Maxon, Contr. U. S. Nat. 
Herb. 10: 487. 1908. (PLATE 2, FIG. 2 and 3) 
Type species, Pteris angustifolia L. (Type specimen from the 
West Indies.) 
Ananthacorus includes a single species, A. angustifolius, which 
has been included during its existence in a large number of genera 
of varying affinities. Goebel pointed out its relationship to the 
Vittarieae on the basis of its spicular cells but did not give it a 
generic name. Several attempts have been made so to signalize it 
but they have failed by reason of nomenclatorial difficulties until 
the action was taken which gave it its present name. Christensen, 
following Diels, included it with Vittaria because of its two spor- 
angial lines (PLATE 2, FIG, 3), but that location can scarcely be 
maintained. These two lines are not homologous with those of 
Vittaria. If they were, they would be located along the costal 
