464 GAGER: CRYPTOMERIC INHERITANCE IN ONAGRA 
The narrow-leaved branch bore flowers and opening buds for 
over a week after the broad-leaved branch had ceased to flower, 
thus pointing to a functional difference. FIGURE 2 shows the 
mature plant just before anthesis began. The greater vigor of 
the broad-leaved branch has obscured the equivalency of the two 
halves, so that the narrow-leaved portion appears in FIGURE 2 to 
be a lateral branch, arising from near the base of the other. Such, 
however, as pointed out above, was not the case. In PLATES 20 
and 21 the morphological features are shown in detail. 
Attempts to secure seed from 15a narrow, either by its own 
pollen or by crossing it with 15a broad, were not successful. The 
broad-leaved branch formed abundant seed when self pollinated, 
and also when crossed with pollen from z5a narrow, but in the 
latter case only one seedling appeared in the seed pan. This 
plant manifested, throughout, only the characters of 15a broad, 
and from it was collected an abundance of close-bred seed, for 
the purpose of testing whether or not a disjunction of characters 
would occur. The F2 generation was grown in the experimental 
garden of the University of Missouri, at Columbia, Mo. Out of 
several hundred seedlings, not one showed the characters of the 
narrow-leaved grandparent, nor did any of these characters ap- 
pear, either by bud sporting or otherwise, in plants that were 
allowed to mature. 
The following explanations of the original anomaly appear 
plausible. 
1. AN INJURY TO THE ANLAGE OF THE PLUMULE. 
2. A QUALITATIVE CHANGE IN EITHER THE EGG OR SPERM that 
united to form the fertilized egg which gave rise to the plant. 
a. In the egg, due to exposure to the radium rays. 
6. In either egg or sperm, spontaneously, or independent of 
the radium. 
3. SECTIONAL BUD-SPORTING IN THE ANLAGE OF THE PLUMULE. 
a. Due to causes resident in the cells (i. €., spontaneous), 
and irrespective of the radium rays. 
(1) The mutation occurred IN THE ANLAGE OF THE PLU- 
MULE. 
(2) The mutation occurred in one of the germ cells, and 
the mutated characters first became active as the 
