410 PENNELL: PLANTS OF SOUTHERN UNITED STATES 
that a number of species may be distinguished upon features at 
first thought of little moment. 
Linnaeus in the first edition of the Species Plantarum recog- 
nized two species, Smilax herbacea and S. Pseudo China. The 
types of these show that they are identical, and that the wide- 
spread glaucous- and glabrous-leaved species of the east is properly 
called S. herbacea. Smilax tamnifolia and S. pulverulenta, two 
surprisingly distinct species considering how poorly our books 
have described them, were added by Michaux. Smilax lasio- 
neuron was described by Hooker in 1840 from the Saskatchewan 
region, and the main innovation of this present revision 1S 
the extension of this species eastward to embrace a plant 
widespread through the Mississippi Valley states. Hooker 
and his followers have laid emphasis upon the short-peduncled 
cymes of S. lasioneuron, but it requires but slight inspection of a 
series of any species of this group to show how undependable 
this character is; indeed, that peduncles should be shortened in 
specimens of western more arid regions seems natural. Two 
southern species have been added by Dr. Small, and the present 
writer describes one species, believed new to science, from Georgia. 
The carrion-flowers are here continued as a subgenus of Smilax, 
in deference to the opinion of Mr. John B. Norton, of the United 
States Department of Agriculture, who is engaged in a study of 
the entire genus Smilax. To him I am indebted for valuable 
criticism. 2 
My interest in this group extends from 1908, and my collections 
include most of the species, although several critical southern 
species have not been obtained. 
Leaves lanceolate to 
iangular-ovate, usually more or less hastate-lobed at base. 
Leaves slightly paler and glabrous beneath. Peduncles 
frequently several in an axil 
Perianth-segments 1.5—2.5 m 
Anthers about equaling the filaments. 
9 
long. Peduncles and pedi- 
cels relatively short, the ce less than 10 mm. long. 1. S. tamnifolia. 
Perianth-segments 4 mm. lo ong. Peduncles and pedicels 
longer, the latter about 20 mm. long 2. S. leptanthera. 
oe shorter than the filaments, 
ot hastate-lobed at base 
IESE leaves pale 
Berri 
“Ridid ovate to cordate, 
badeatk relatively short-acuminate. 
es dark-blue, glaucous. Peri anth-segments 
broadly lanceolate, as mm. lor 
