SAX: FERTILIZATION IN FRITILLARIA PUDICA 515 
Both polar nuclei appear to be in a resting condition. The male 
nucleus may be attached on any side of the upper polar nucleus, 
The cases of migration of the upper polar nucleus together 
with the male nucleus were not found fre- 
quently, only forty-seven cases being found 
in the great number of fertilization stages. 
However, the cases of triple fusion of the 
polar nuclei and male nucleus were very 
abundant. 
TEXT FIG. 3 shows the embryo sac in 
longitudinal section with a synergid, the egg 
and male nucleus in contact, and the two 
polar nuclei and second male nucleus in con- 
tact. The male nucleus stains darker and is 
between the polar nuclei. It will be noted 
that there is very little cytoplasm in the 
upper end of the ‘‘Endospermanlage”’ at this 
time, which is very characteristic. It might 
appear, on comparing TEXT FIGS. 2 and 3, 
that the lower polar nucleus migrates also to 
meet the two approaching nuclei, but if there 
is a migration of the lower polar nucleus, it 
is slight, as has been observed in numerous 
examples, and it is accidental that the two 
sacs drawn should differ so markedly in this 
respect. 
Fics. 33 and 34 show triple fusion. In 
Fic. 34, the male nucleus is at the side, while 
in Fic. 33, it is between the two polar nuclei. a 
At this stage the chromatin of the male nu- jucteus; the other male 
cleus has lost some of its net-like structure nucleus and upper polar 
and contains large pieces of chromatin with rt ie ipa seni 08 
delicate threads, and in some cases, nucleoli. 
FIG. 32 shows a case where the two polar nuclei and male nucleus 
are in contact, but the latter is still vermiform and its contents 
are in a net-like structure. 
The polar nuclei remain in contact without fusing for con- 
siderable time. At the time of fusion the chromatin contents 
