204 Evans: NOTES ON GENUS HERBERTA 
The perianth of Herberta, when well developed, clearly supports 
the theory that three floral leaves take part in the formation of this 
organ (PLATE 8, FIG. 13). Itissplit for nearly half its length into six 
slender laciniae, essentially like divisions of leaves, every alternate 
split being a little deeper than the others. These deeper splits mark 
the boundaries of the three coalescent leaves. In the undivided 
portion of the perianth a cross section shows that six rounded 
ridges are present, separated by six rounded grooves (PLATE 8, 
FIG. 14). Three alternate grooves are deeper than the others and 
correspond with the three deeper splits at the mouth of the 
perianth. The innermost bracts and bracteoles, each with its 
two dorsal ridges separated by a groove, are closely appressed to 
the ridges and grooves of the perianth. The laciniae of the 
perianth bear numerous slime-papillae and often teeth. The slime 
papillae occur not only along the margins of the laciniae toward 
the base but also on the inner surface of the perianth. Here they 
sometimes form short rows and sometimes show no regular ar- 
rangement. The surface papillae are occasionally borne on the- 
tips of short teeth or along the margins of narrow and irregular 
ridges. In some cases the perianth is not developed normally. 
More than six laciniae, for example, may be present at the mouth, 
or one of the splits may extend to the very base; sometimes both 
of these abnormalities may be seen in a single perianth. An 
increase in the number of laciniae is not surprising, since ordinary 
vegetative leaves with three divisions sometimes occur, and the 
deep split might plausibly be interpreted as a place where coa- 
lescence had failed to take place. 3 
The wall of the capsule is described in its essential features by 
Spruce. It is composed of from five to seven layers of cells 
(TEXT FIG. 9) and shows a thickness of 70-100 u. This is con- 
siderably more than in most of the leafy Hepaticae studied by 
Andreas.* In Plagiochila asplenioides (L.) Dumort., for example, 
where the wall is composed of seven or eight layers, the thickness 
is only 55 u, while in Chiloscyphus polyanthus (L.) Corda, where 
the wall is composed of five layers, the thickness is only 25 4. It 
is less, however, in Herberta than in Pleurozia purpurea (Lightf.) 
* Ueber den Bau der Wand und die Oeffnungsweise des Lebermoossporogons- 
Flora 86: 161-213. pl. r2+f. 1-25. 1899. 
