334 BERRY: A MIDDLE EOCENE GONIOPTERIS 
clays of the Yegua formation at Columbia, Louisiana, occurring 
also in the sandy clays of the Lisbon formation near Newton, 
Mississippi, but none of the collected material is in fruit. The 
form and venation are so well marked and distinctive, however, 
that the species is at once correlated with the rather abundant 
Tertiary type that, under the name of Goniopteris, Lastrea or 
Phegopteris, is so characteristic of the fern floras of that time. It 
is at once distinguished from all of these by the peculiar venation, 
otherwise it shows the same habit, winged rachis and outline as, 
for example, Lastrea stiriaca Heer* of the European Oligocene, 
described originally by Ungerf as Polypodites, referred to Gontop- 
teris by Alexander Braunt and to Phegopteris by Ettingshausen.§ 
There are at least fifteen known Tertiary species of this general 
type, most of which are European, although several have been 
recorded from American localities. Most of the foreign material 
is somewhat younger than the present species, although two dif- 
ferent forms have been described from the middle Bagshot beds 
of southern England, a nearly homotaxial (Lutetian) horizon. 
Several early Eocene species have been recorded in this country 
from the Rocky Mountain region. In addition to the differences 
in venation previously referred to, Lastrea intermedia Lesquereux|| 
from the Denver formation has the pinnae decurrent on the main 
stipe; Lastrea Goldiana Lesquereux" from the same horizon has 
crenulate and deeply divided margins and simple tertiaries; the 
form from Sand Creek, Colorado, referred to Ettingshausen’s 
Monte Promina species Lastrea polypodioides**, has a denticulate 
margin and simple tertiaries. The form from the lower Eocene 
of Oregon identified by Newberry as Lastrea Knightianat} and 
commonly referred to the European early Miocene species Lastrea 
Fischert Heer{{ is much like the present species in size and general 
* Heer, Fl. Tert. Helv. 1: 31. pl. 7, 8. oe 3: 151. dl. 143. 
ft Unger, Chlor. Protog. rat. pl. 36. 
}¢ Braun, Zeits. Deutsch. Geol. Gese 4: ee 1852 
§ Ettingshausen, Foss. FI. Bilin. 1: 16. pl. 2, f. ie. 1866. 
|| Lesquereux, Tertiary Flora, 56. pl. 4, f. 14. 1878. 
{ Idem, f. 13. 
** Idem, 57. pl. 4, f. II, 12. y 
tt Newberry, Proc. U. S. Nat. Mus. 5: 503. 1882. 
tt Heer, Fl. Tert. Helv. 1: 34. pl. 0, f. 3. 1855; Lesquereux, Cret. & Tert. Fl. 239- 
pl. 50, f. 1, ra. 1883; Newberry, Mon. U. S. Geol. Surv. 35: 10. pl. 48, f. 6. 1898. 
1859. 
