502 ARTHUR: RELATIONSHIP OF GENUS KUEHNEOLA 
called K. Uredinis (Link) Arth., if the earliest specific name is 
to be adopted (which happens to be one founded upon the telia), 
is now known to occur on many species of Rubus throughout 
Europe and North America, and another species, K. andicola 
(Diet. & Neg.) Diet., is found in South America on Rubus geoides 
Sm. In 1912 Dietel and the writer added three species to the 
genus on other rosaceous hosts, and between 1912 and 1914 
Butler, Sydow and the writer added eight other species, whose 
hosts belong to the families Vitaceae, Artocarpaceae, Burseraceae, 
Anacardiaceae, Verbenaceae and Bignoniaceae. Of all these 
species the full life history is known for only the first species, 
K. Uredinis on Rubus (Rosaceae), and the last one, K. Markhamiae 
(P. Henn.) Syd. on Markhamia (Bignoniaceae). 
The last species occurs in German East Africa. A good 
description is given in Sydow’s Monographia Uredinearum (3: 
318), but without mentioning the pycnia. These were found on 
an original specimen in my own herbarium. ‘They are epiphyllous, 
in small groups opposite the primary uredinia, punctiform, sub- 
cuticular, 100-110 u in diameter, the hymenium flat; ostiolar 
filaments wanting. The urediniospores are borne singly on pedi- 
cels, sparsely echinulate-verrucose, and with three equatorial pores. 
It will be seen that the general characters of K. Uredinis and 
K. Markhamiae coincide, and the two species can be considered 
genuine representatives of the genus Kuehneola. Although the 
pycnia of K. andicola have not been detected, yet the similarity 
of the known characters, and the host being a species of Rubus, 
that species can also be included in the genus without hesitancy. 
The other recorded species on rosaceous hosts are K. japonica 
Diet., on three species of Rosa in Japan, K. Duchesneae Arth., 
on Duchesnea indica (Andr.) Focke (Fragaria indica Andr.), in 
North America, and K. obtusa (Strauss) Arth., on three or more 
species of Potentilla, in Europe, and on P. canadensis L. in North 
America. No specimen of the species on Rosa has been seen by the 
writer, but the description, for telia only, appears to warrant its 
inclusion under Kuehneola. 
In studying the species on Potentilla, or two species as main- 
tained by some uredinologists, it was brought out some time since 
that the relationship with species of Kuehneola on Rubus is not 
