﻿146 
  Harper: 
  Inheritance 
  of 
  sugar 
  

  

  + 
  

  

  may 
  be 
  areas 
  on 
  a 
  few 
  seeds 
  which 
  closely 
  resemble 
  the 
  wrinkled 
  

   condition 
  of 
  sweet 
  seeds. 
  In 
  general 
  it 
  is 
  stated 
  this 
  pseudo- 
  

   starchy 
  race 
  when 
  crossed 
  with 
  pure 
  starchy 
  races 
  gives 
  in 
  the 
  Fg 
  

   segregation 
  into 
  sugary 
  and 
  starchy 
  kernels 
  as 
  if 
  it 
  had 
  been 
  a 
  

   so-called 
  pure 
  sweet 
  though 
  the 
  F2 
  kernels 
  so 
  obtained 
  show 
  con- 
  

   siderable 
  tendency 
  to 
  pseudo-starchiness 
  in 
  later 
  generations 
  in 
  a 
  

   fashion 
  that 
  makes 
  the 
  results 
  hard 
  to 
  classify. 
  The 
  explanation 
  

   of 
  his 
  observations, 
  Jones 
  believes, 
  is 
  to 
  be 
  sought 
  in 
  the 
  recog- 
  

   nition 
  that 
  more 
  factors 
  are 
  concerned 
  in 
  the 
  determination 
  of 
  the 
  

  

  r 
  

  

  sweet 
  and 
  starch 
  pair 
  of 
  characters 
  than 
  had 
  before 
  been 
  realized. 
  

  

  He 
  makes 
  no 
  claim 
  to 
  have 
  reached 
  an 
  adequate 
  or 
  theoretically 
  

   satisfying 
  analysis 
  from 
  the 
  Mendellan 
  standpoint. 
  His 
  assumed 
  

   three 
  factors 
  are 
  merely 
  taken 
  "as 
  an 
  illustration" 
  and 
  he 
  speaks 
  

   of 
  the 
  "indefinite 
  nature 
  of 
  the 
  character/' 
  

  

  Jones 
  refers 
  (p. 
  388) 
  to 
  other 
  cases 
  he 
  has 
  observed 
  in 
  which 
  

   intermediate 
  kernels 
  have 
  appeared 
  in 
  starch 
  and 
  sugar 
  corn 
  

   crosses, 
  though 
  he 
  expresses 
  no 
  opinion 
  as 
  to 
  the 
  frequency 
  of 
  

   their 
  occurrence, 
  and 
  leaves 
  the 
  question 
  of 
  their 
  nature 
  and 
  

   origin 
  much 
  as 
  it 
  was 
  before. 
  He 
  recognizes 
  the 
  possible 
  influence 
  

   of 
  the 
  fact 
  that 
  the 
  mother 
  plant 
  is 
  always 
  one 
  generation 
  in 
  

  

  r 
  

  

  advance 
  of 
  the 
  endosperm, 
  though 
  a 
  real 
  test 
  of 
  the 
  significance 
  

   of 
  this 
  situation 
  is 
  difficult 
  to 
  achieve. 
  

  

  On 
  the 
  assumption 
  of 
  relatively 
  fixed 
  unit 
  factors, 
  it 
  is 
  rather 
  

   striking 
  when 
  this 
  intermediate 
  condition 
  has 
  appeared 
  that 
  new 
  

   and 
  definite 
  combinations 
  of 
  the 
  factors 
  assumed 
  are 
  so 
  difficult 
  to 
  

   achieve. 
  Neither 
  the 
  pseudo 
  starchy 
  nor 
  the 
  segregated 
  sweet, 
  

   either 
  in 
  appearance 
  or 
  behavior, 
  when 
  bred 
  seem 
  quite 
  to 
  meet 
  

   the 
  standards 
  of 
  a 
  homozygous 
  race 
  in 
  the 
  strict 
  sense 
  of 
  Johannsen. 
  

   We 
  seem 
  to 
  have 
  here 
  the 
  familiar 
  phenomenon 
  which 
  was 
  known 
  

   to 
  the 
  older 
  breeders 
  as 
  "breaking 
  up" 
  of 
  types 
  in 
  the 
  second 
  

   and 
  following 
  generations 
  as 
  a 
  result 
  of 
  hybridizing- 
  From 
  the 
  

   standpoint 
  of 
  current 
  theories 
  we 
  are 
  inclined 
  to 
  focus 
  our 
  attention 
  

  

  w 
  

  

  upon 
  the 
  possibility 
  of 
  recovering 
  parent 
  types 
  after 
  hybridizing 
  

   while 
  the 
  older 
  breeders 
  regarded 
  this 
  phenomenon, 
  though 
  

   familiar, 
  as 
  of 
  little 
  Interest 
  compared 
  with 
  the 
  cases 
  in 
  which 
  

   "breaking 
  up" 
  occurred. 
  The 
  search 
  was 
  frequently 
  so 
  far 
  as 
  

   hybridization 
  was 
  used 
  in 
  breeding 
  for 
  practical 
  ends 
  for 
  a 
  cross 
  in 
  

   which 
  breaking 
  up 
  would 
  occur. 
  A 
  species 
  with 
  desirable 
  basic 
  

  

  