﻿170 
  Harper: 
  Inheritance 
  of 
  sugar 
  

  

  evidence 
  that 
  any 
  such 
  assumed 
  factors 
  are 
  distributed 
  In 
  the 
  

   reduction 
  divisions 
  in 
  any 
  clear-cut 
  and 
  obvious 
  fashion, 
  such 
  as 
  

   the 
  simple 
  mono-hybrid 
  ratios, 
  and 
  to 
  name 
  in 
  the 
  germ 
  plasm 
  

   the 
  same 
  qualities 
  observed 
  in 
  the 
  soma 
  explains 
  nothing 
  and 
  is 
  

  

  m 
  

  

  merely 
  the 
  old 
  preformationism 
  in 
  a 
  modern 
  guise. 
  

  

  Correns 
  ('oi) 
  and 
  East 
  and 
  Hayes 
  ('ii) 
  have 
  based 
  their 
  

   generalization 
  that 
  the 
  qualities 
  starchy 
  and 
  sugary 
  make 
  a 
  

   typical 
  Mendelian 
  pair 
  of 
  characters 
  in 
  their 
  segregation 
  and 
  

   recombination 
  behavior 
  on 
  the 
  assumption 
  that 
  intermediates 
  

   behave, 
  when 
  grown 
  and 
  selfed, 
  practically 
  like 
  the 
  visibly 
  more 
  

   typical 
  starchy 
  or 
  sweet 
  kernels, 
  except 
  in 
  rare 
  cases 
  of 
  mutation, 
  

   when 
  as 
  East 
  reports 
  an 
  occasional 
  intermediate 
  ear 
  (i 
  in 
  10,000) 
  

   appears 
  with 
  no 
  evidence 
  that 
  it 
  is 
  a 
  result 
  of 
  hybridization 
  and 
  

   whose 
  offspring 
  tend 
  to 
  breed 
  true, 
  thus 
  causing 
  trouble 
  to 
  growers 
  

  

  m 
  

  

  J 
  

  

  of 
  sweet 
  corn 
  for 
  canneries 
  in 
  case 
  such 
  ears 
  are 
  not 
  avoided 
  as 
  

   seed. 
  

  

  That 
  however 
  both 
  in 
  their 
  behavior 
  in 
  the 
  following 
  gener- 
  

   ation 
  and 
  in 
  their 
  appearance 
  there 
  is 
  adequate 
  ground 
  for 
  treating 
  

   the 
  intermediates 
  as 
  a 
  special 
  class 
  is, 
  it 
  seems 
  to 
  me, 
  sufficiently 
  

   evident 
  from 
  the 
  data 
  given 
  above. 
  In 
  Table 
  II, 
  section 
  i, 
  D, 
  I 
  

   have 
  brought 
  together 
  and 
  compared 
  the 
  results 
  obtained 
  by 
  

   planting 
  intermediates 
  with 
  those 
  obtained 
  by 
  planting 
  typical 
  

   starchy 
  and 
  sugary 
  kernels 
  from 
  ear 
  130 
  d. 
  About 
  40 
  per 
  

   cent 
  of 
  intermediates 
  were 
  obtained 
  from 
  planting 
  intermedi- 
  

   ates, 
  while 
  ears 
  from 
  sweet 
  and 
  starchy 
  kernels 
  gave 
  only 
  about 
  

   5 
  per 
  cent 
  of 
  intermediate 
  kernels. 
  The 
  evidence 
  from 
  Correns's 
  

   work 
  as 
  well 
  as 
  my 
  -own 
  is 
  overwhelming 
  that 
  such 
  intermediate 
  

   kernels 
  do 
  occur 
  sporadically 
  but 
  quite 
  commonly 
  as 
  a 
  result 
  of 
  

   crossing 
  as 
  well 
  as 
  congenitally 
  as 
  a 
  result 
  of 
  sexual 
  reproduction 
  

   in 
  so-called 
  pure 
  races 
  and 
  that 
  in 
  whatever 
  way 
  they 
  arise, 
  they 
  

   do 
  not 
  at 
  once 
  breed 
  true. 
  By 
  selection 
  the 
  percentage 
  of 
  such 
  

   kernels 
  may 
  be 
  increased 
  with 
  fair 
  probability 
  in 
  every 
  case 
  that 
  

   rather 
  pure 
  races 
  of 
  intermediates 
  may 
  be 
  established. 
  Selection 
  

   of 
  intermediates 
  through 
  the 
  F2 
  and 
  F3 
  generations 
  selfed 
  followed 
  

   by 
  an 
  out 
  cross 
  with 
  an 
  intermediate 
  of 
  different 
  parentage 
  gave 
  

   as 
  the 
  progeny 
  of 
  one 
  ear 
  (166 
  a) 
  three 
  out 
  of 
  four 
  ears 
  with 
  a 
  

   considerable 
  excess 
  of 
  intermediates 
  and 
  with 
  no 
  pure 
  starchy 
  

   kernels. 
  The 
  fourth 
  ear 
  was 
  an 
  almost 
  pure 
  sweet. 
  The 
  off- 
  

  

  