DopGE: RELATIONSHIPS OF FLORIDEAE AND ASCOMYCETES 161 
'ptivatim unterrichtet bin, war sowohl De Bary wie Schmitz 
dieser Annahme zugeneigt.”’ 
Woronin, Janczewski, De Bary, and Borzi do not specifically 
identify the tapering end of the ascogonium in Ascobolus as a 
trichogyne though they give no other interpretation of this very 
important structural feature of the egg apparatus, and Sachs’ 
classic figure of A. furfuraceus represents the fertilization as 
occurring by means of a pollinodium which applies itself to the tip 
of the many-celled ascogonium, though this conception has by 
no means been held by all subsequent students of the form. 
Stahl’s work on the lichens established beyond question the exis- 
tence of trichogynes fertilized by free spermatia, and this is prob- 
ably the most important discovery bearing on the relationship 
of the two groups. 
Among the Ascomycetes the trichogyne is most characteris- 
tically developed in the Laboulbenieae. The variability in the 
form of the trichogynes in this group is almost without limit. 
They may be one-celled structures as in Stigmatomyces Baeri 
(F IG. 2, G), recalling the trichogyne of Batrachospermum (FIG. 4, 
B), or they may be multicellular spirally coiled and branched as 
in Compsomyces verticillatus (F1G. 2, 1). Some of them are very 
similar to the trichogyne of Pyronema (Fic. 2, D),as in Zodiomyces 
vorticellarius (FIG. 2, J), where the spermatia remain attached to 
the stalk, and the trichogyne is therefore a structure active in 
seeking out the spermatia. 
Ascodesmis, a form closely related to Pyronema, furnishes a 
further example of a reduced trichogyne. Claussen (25) finds 
that the end cell of the short spiral coil of the ascogonium functions 
as a trichogyne (Fic. 2,E). It is not especially different in struc- 
ture from the other cells of the ascogonium. 
Kihlman (57) did not consider the tapering end of the asco- 
gonium of Melanospora as a trichogyne (Fic. 2, K) for the reason 
that he was unable to find that it fused with any hypha repre- 
senting an antheridium. Miss Nichols (64), however, reports in 
Hypocopra and Ceratostoma (Fic. 2, C), other members of the same 
Sroup, that the end cell of the ascogonium does connect with an 
antheridium. 
Fraser and Chambers (46) find that the ascogonium of A sper- 
