162 DopGE: RELATIONSHIPS OF FLORIDEAE AND ASCOMYCETES 
gillus herbariorum is provided with an end cell that functions asa 
trichogyne in certain cases. They think this form is becoming 
apogamous, but this fact would not, in their opinion, affect the 
question of the homologies of the cells at the end of the archicarp. 
I have described a form of archicarp in Ascobolus carbonarius 
which in its origin is entirely unlike any hitherto known in the 
Ascomycetes. I refer to the common form of the species which 
produces abundant spore-like bodies, ‘‘conidia,’’ on long stalks, 
and not to the form A. mirabilis (37) of Dangeard. This latter 
strain which I have since isolated and grown several months in 
pure cultures produces the ascogonium directly from the mycelial 
hyphae only rarely producing conidia, and then only in very old 
cultures. Other interesting abnormalities in the archicarp crop 
out frequently in this strain. I have not been able to follow the 
development of these abnormal forms but they do not seem to give 
rise to normal apothecia. 
The typical ascogonia of A. carbonarius develop apothecia in 
about two weeks. As described (37), most frequently the asco- 
gonium arises directly from what appears to be an asexual conidium 
which germinates forming a coiled portion. The original germ 
tube thus becomes the stalk of the archicarp at the end of which 
is formed a second coil of extremely large cells composing the 
ascogonium proper, and from this is derived by sudden constric- 
tion of the end cell a long irregularly coiled trichogyne (FI. 1, E) 
which in general is not very unlike the trichogyne of Collema 
pulposum (Fic. 1,C) described by Miss Bachmann (2, 3). When 
grown in artificial media the trichogyne winds about in the 
medium or on its surface somewhat irregularly and in many 
cases the long end cell comes in contact with other conidia on long 
stalks, which I have called antheridial conidia (Fics. 1, E, and 3, 
B). The trichogyne end coils itself tightly about this antheridial 
conidium in such a manner as to indicate that there is a definite 
attraction between them. The resemblance of the antheridial 
conidia here to the spermatia of Miss Bachmann’s Collema (FIGS. 
1, C, and 3, D) is direct and striking. I do not know that any of 
these conidia of A. carbonarius function as asexual bodies in 
distributing the species. They do not become detached from their 
stalks and are not oriented as we should expect functional conidia 
