DopGE: RELATIONSHIPS OF FLORIDEAE AND ASCOMYCETES 163 
tobe. The conidia of Lachnea abundans are very loosely attached 
to erect aerial conidiophores and germination takes place as soon 
as they are brought into a fresh medium. In my cultures these 
conidia of A. carbonarius never function in any other way than 
that described. They never produce mycelia directly. Aborting 
ascogonia do sometimes grow out vegetatively into long hyphae 
but such growths are recognizable at once and would not be 
mistaken for trichogynes. The trichogyne is a hypha of very 
uniform diameter terminating rather bluntly and without branches 
(Fic. 1, E). 
G. I. Septate trichogynes of Ascomycetes. A. Collema microphyllum; B. 
Physma compactum; C. Collema pulposum; D. Polystigma rubrum; E. Ascobolus 
carbonarius; F. Lachnea cretea; G. Lachnea melaloma; H. Ascobolus furfuraceus. 
45g, ascogonium; an.c, antheridial conidium; p, pore in cross wall; pd, callous pad; 
SPE, spermogonium; st.h, sterile hyphae; stk, stalk of ascogonium; fr, trichogyne. 
A, B, Stahl; C, Bachmann; D, Fisch; F, Fraser; H, Harper; E, G, original. 
The importance of Miss Bachmann’s work on Collema pulposum 
can scarcely be overestimated since it furnishes perhaps the most 
Convincing evidence in support of the view that the spermatia of 
the lichens are sexual organs and that their trichogynes are 
morphologically equivalent to those of the red algae and of Pyro- 
