166 DopcGE: RELATIONSHIPS OF FLORIDEAE ‘AND ASCOMYCETES 
Lecanora subfusca, and Harper (53) figures a two-branched 
trichogyne in Pyronema. Miss Bachmann (3) also describes and 
figures a branch which is given off from the ascogonium at least 
very close to the base of the trichogyne in Collema pulposum. 
The Tulasnes (90) mention the existence of an ascogonium in 
Lachnea melaloma. 1 have grown this form abundantly in cul- 
tures on agar media and find an archicarp of striking proportions. 
The long stalk, the many-celled coiled ascogonium prolonged into 
a trichogyne-like portion (Fic. 1, G), likewise the great irregularity 
and variety of forms which it shows in this species suggest a 
relationship with Ascobolus carbonarius. It may be even more 
Fic. 3. Antheridia and spermatia. A. Corallina officinalis; B. Delesseria 
sanguinea; C. Collema microphyllum; D. Collema pulposum; E. Ascobolus magnificus; 
F. Pyronema; G. Sphaerotheca; H. Ascobolus carbonarius. an, antheridium; 4.6, 
antheridial conidium; spm, spermatium; stk, stalk. A, Guignard; B, Svedelius; 
C, Stahl; D, Bachmann; F, Tulasne; G, De Bary; E, H, original. 
closely related to the strain A. mirabilis since it produces no conidia, 
chlamydospores, or any asexual spore bodies whatever. The 
Tulasnes (go) figured conidia for Pyronema; later (89) they re- 
ferred them to Lachnea melaloma. They undoubtedly were mis- 
taken in both cases, as I find no such spores in my cultures. The 
archicarp of Lachnea melaloma frequently branches either in the 
region representing the trichogyne, or well back near the basal 
portion. In the latter case the branching is somewhat dichoto- 
mous. There is also a marked tendency to dichotomous branching 
in the vegetative hyphae. 
As compared with the trichogynes of the Ascomycetes in 
