DopGE: RELATIONSHIPS OF FLORIDEAE AND ASCOMYCETES 175 
thallus through which the trichogyne penetrates to the surface 
(Fic. 4, F). The auxiliary cell is derived from the lower cell of 
one of the sterile filaments and is brought into connection with the 
carpogonium by a short odblastema. Daines was unable to dis- 
cover that this filament united with any other cell as in the case 
of other species of the Grateloupiaceae described by Berthold, 
and since procarpic branches and auxiliary cells are borne in pairs 
Daines concludes that this species should be placed with the 
Ceramiales. 
+ 5. Trichogynes aes cnayar ae ~ the carpogonium in ‘ie po algae. 
A, ht Pe atin nee m; C, D. Dudresnaya purpurifera; E. Dudresnaya 
coccinea. aux, auxiliary cell; Ee br, carpogenic branch; cpg, carpogonium; o.f, 
odblastema filament; tr, trichogyne. A, B, D, E, Oltmanns; C, Schmitz. 
Vastly more complex and perhaps furnishing still more evi- 
dence as to the ancestry of the Ascomycetes are the Cryptone- 
miales, the forms upon which Schmitz based his theory of sec- 
ondary sexual fusions. 
The procarpic branch in Cruoriopsis cruciata (Schmitz, 78, 
80) is imbedded within the thallus but its trichogyne extends out 
beyond the surface. Near these procarpic branches other fila- 
ments are developed which Schmitz called ‘‘procarps without 
trichogynes.” The end cells (auxiliary cells) of these branches 
remain imbedded in the thallus and do not grow out into tricho- 
Synes. After the fertilization of the procarp there now grows out 
from its under part an odblastema filament which branches freely 
and spreads widely inside of the thallus and “‘fertilizes the procarp 
