490 RYDBERG: NOTES ON ROSACEAE 
retained up to 1906, when Dr. Greene segregated from it Acomasty- 
lis and Erythrocoma. In my opinion the former was rightly 
taken out, as it differs from Sieversia by the same character of 
the fruit that separates Anemone from Pulsatilla. Erythrocoma 
on the other hand I can not regard as distinct generically from 
Sieversia. Its species differ from the type of Sieversia only in 
the erect instead of spreading petals and a better developed 
hypanthium. If Erythrocoma should be kept distinct, Gewm rivale 
should: be taken out of Geum, as it differs by just the same char- 
acters, and still that species frequently hybridizes with the other 
species. Dr. Greene made the following statements which are 
not exactly true: ‘‘Sieversia was founded on a Siberian under- 
shrub, low and slender, with almost rotate calyx and corolla, the 
former nearly chorisepalous, and its mature styles are long, filiform, 
plumose to the very apex.” There are apparently three species 
which have gone under the name Sieversia anemonoides, of which 
one can be called an undershrub, the other two having the habit of 
Dryas octopetala, being strongly cespitose with the branches more 
or less covered by soil and moss. I think the type of Sieversia 
was one of these. The difference between this cespitose stem and 
the branching rootstock of Erythrocoma is indeed very small. 
The styles of Sieversia anemonoides Willd., or S. pentapetala (L.) 
Greene, are not plumose to the apex, for the upper part is naked, 
soft, withering, as it is in Erythrocoma, the only difference being 
that it is very short, scarcely 2 mm. long. This naked portion 
of the style is found not only in the type of Sieversia and in S. 
ciliata and its relatives, but also in the S. montana group, to which 
S. radicata, S. Peckii and S. calthifolia belong. The naked portion 
in most is rather persistent, although usually withering in age, 
but in some species or even individuals it breaks off. It is how- 
ever never articulated to the lower portion of the style as in Geum, 
where the upper portion, which is usually hairy, falls off very 
early. C. P. Smith* describes the style of Erythrocoma ciliata as 
follows: “Style hooked, or articulated [italics mine], the terminal 
portion often deciduous.” I have seen it somewhat bent or even 
occasionally hooked, but never with a proper articulation. The 
soft upper naked portion, after it has withered, often falls off at 
* Muhlenbergia 8: 7. 1912. 
ee eer 
