1920.] G. A. BouLENGER : Frogs of the Genus Rana. 65 



obliquely directed upwards, the upper eyelid often verj' narrow ; 

 nostril a little nearer the end of the snout than the eye ; the dis- 

 tance between the nostrils not or but little greater than the inter- 

 orbital width, which usually nearly equals or (in large males) exceeds 

 that of the upper eyelid ; tympanum usually hidden, rarely slightly 

 distinct, J to J the diameter of the eye and widely separated from 

 the latter. 



Fingers short or moderate, obtuse or slightly swollen at the 

 end, first and second equal or first the longer, third as long as or 

 longer than the snout, second and third often with a dermal fold on 

 each side; subarticular tubercles large, moderately prominent. 



Hind limb short or rather short, the tibio-tarsal articulation 

 reaching the temple or the eye, the heels meeting or failing to 

 meet, when the limbs are folded at right angles to the body ; 

 tibia 2j to 3 times as long as broad, 2 to 2^ times in length from 

 snout to vent, as long as or a little shorter than the foot, shorter 

 than the fore limb. Toes short or rather short, the tips dilated into 

 small 1)ut very distinct discs, iuWy webbed or the web strongh- 

 emarginate and not reaching beyond the penultimate phalanx of 

 the fourth ; subarticular tubercles rather large, moderately promi- 

 nent ; a narrow tarsal fold; inner metatarsal tubercle narrow, 

 moderately prominent, I to t the length of the inner toe ; no outer 

 tubercle. 



Skin smooth above, except on the calves and on the posterior 

 part of the upper eyelids, which are nearly always warty, or with 

 short longitudinal glandular folds or roundish tubercles ; a glandu- 

 lar fold from the eye to the shoulder ; sometimes a distinct fold 

 across the head, behind the eyes. Lower parts smooth. 



Brown above, uniform or with small darker spots on the back 

 and more or less irregular cross-bars on the limbs ; usuall}^ a dark 

 bar, light-edged in front, between the ej-es, and dark vertical bars 

 on the lips ; a dark canthal and temporal streak often j^resent ; a 

 yellow vertebral streak rarel}' present (specimens from Matang, 

 Bongo, Kina Balu, and Kuatun). Lower parts white, throat and 

 breast often mottled with brown, sides of belly and lower surface 

 of legs sometimes speckled with brown. 



Males without vocal sacs, the fore limbs neither thickened 

 nor bearing nuptial excrescences ; differing from the female by the 

 larger, often much larger, head, which may be more triangular in 

 shape, and by the larger bony prominences in front of the lower 

 jaw. 



Nasals large, in contact with each other and with the fronto- 

 parietals, which cover the ethmoid ; zygomatic branch of squa- 

 mosal very long, curved, reaching the lower border of the eye. 

 Omosternal style slender, forked at the base; sternal style short. 

 Terminal phalanges transversely expanded at the end. 



The vitellus of eggs ready to be laid measures 2 millim. in dia- 

 meter, the mother measuring 51 millim. from snout to vent. 



Tadpoles from Hong Kong Peak, referred by Annandale to 

 R. kuhlii^ and said to resemble those of R. macrodon, have since 



