1913. | S. Kemp: Crustacea Stomatopoda of the Indo-Pacific Region. 145 
Genus Gonodactylus, Latreille. 
1825. Gonodactylus, Latreille, Encycl. Méthod., X, p. 473. 
1837. Gonodactylus, H. Milne-Edwards, Hist. Nat. Crust., II, p. 528 (partum). 
1849. Gonodactylus, De Haan, in Siebold’s Fauna Japonica, Crust., p. 220. 
1880. Gonodactylus, Miers, Ann. Mag. Nat. Hist. (5), V, p. 115 (partim). 
1886. Gonodactylus, Brooks, Voy. H.M.S. ‘ Challenger,’ XVI, Stomatop., p. 55. 
1886. Protosquilla, Brooks, tbid., p. 65. 
1894. Gonodactylus, Bigelow, Proc. U.S. Nat. Mus., XVII, p. 492. 
1894. Protosquilla, Bigelow, ibid., p. 492. 
1895. Gonodactylus, Hansen, Isop. Cumac. u. Stomatop., d. Plankton-Exped., p. 86. 
1906. Gonodactylus, Nobili, Ann. Sci. Nat. Zool. (9), IV, p. 326. 
Carapace little, if at all, narrowed anteriorly ; without carinae; gastric grooves 
distinct, cervical groove wholly absent; antero-lateral angles rounded or acute. 
Cornea of eye sometimes indistinctly bilobed, mandibular palp present and composed 
of two or threesegmentsorabsent. Ischio-meral articulation of raptorial claw situated 
at a point in advance of proximal end of merus, the latter segment consequently 
extending backwards beyond the joint ; inferior surface of merus not channelled longi- 
tudinally throughout its entire length; upper margin of propodus not pectinate ; 
dactylus inflated at base, without teeth on inner margin. Shorter ramus of last three 
thoracic appendages linear, composed of two segments. Free thoracic and abdominal 
somites compressed, no sharp carinae on first four abdominal somites. Telson very 
variable in form and sculpture, carinate, tuberculate or spinous. Ventral process of 
uropods terminating in two sharp spines, inner usually shorter than outer. 
There are no well-marked secondary sexual distinctions. 
In reducing Brooks’ Protosquilla to a synonym of Gonodactylus I have followed 
Hansen and Nobili. The character which Brooks employed, the fusion or free articula- 
tion of the last abdominal somite and telson, is, as Hansen has shown, wholly unreliable 
and no other feature sufficiently well-marked to serve as a generic distinction can be 
found. On the contrary it seems that the species here included in Gonodactylus fall 
into four natural groups of approximately equal value, but these do not, I believe, possess 
any greater claims to nomenclatorial distinction than do the groups in Lystosquilla 
or in other genera. 
The four groups may be distinguished by the following characters :— 
Group I.—The anterior margin of the carapace slopes forward on either side of the rostrum, so that 
the antero-lateral angles are well in advance of the rostral base. ‘The rostrum has a sharp median spine and 
its antero-lateral angles are rounded or subacute : in rare cases acute. ‘The mandibular palp is composed 
of three distinct segments. The posterior margin of the sixth abdominal somite is straight or slightly 
convex in dorsal view. ‘The basal segment of the outer uropod projects posteriorly a little beyond the 
articulation of the ultimate segment, and its external margin is armed with a series of straight, or almost 
straight, movable spines. The inner uropod is normal in shape and, asa rule, in the disposition of its setae. 
This group includes G. chivagra, demani, graphurus and their immediate allies. 
Group II.—The anterior margins of the carapace slope forwards as in the preceding group. The 
rostrum is trispinous: the antero-lateral angles are produced to sharp spines often almost as slender as the 
