1913.] S. Kemp: Crustacea Stomatopoda of the Indo-Pacific Region. 153 
But G. spinosus, which is unquestionably more remote from G. chivagra, he includes 
under the latter name as a variety. 
G. demanti is, it seems, a smaller form than G. chiragra and does not exceed 40 mm. 
in length. ‘The spinulation of the telson is most astonishingly variable (as will be seen 
by reference to pl. ix, figs. rog-111) ; but even if, as is not unlikely, an individual 
wholly destitute of spinules should appear, the other characters noted above will, I 
believe, remain amply sufficient to justify specific separation from G. chiragra. The 
variation in the number of spinules is unquestionably continuous. 
Gonodactylus spinosus, Bigelow, described one month later than Henderson’s 
G. demani, differs from that species only in the slightly greater proportional length of 
the telson and in the suppression of the intermediate teeth of the margin. But G. 
demani itself shows no little variation in this respect, and in the Indian specimens 
which I have referred to spinosus, these teeth are slightly better developed than in the 
typical individual figured by Lanchester. I am forced therefore to the conclusion, 
already suggested by Nobili, 1906 (a), p. 331, that G. spinosus can, at most, only be 
retained as a variety of G. demani.'! The form appears to be rare ; in the details of its 
spinulation it is perhaps as variable as G. demani,' but on this point little is known. 
Borradaile’s Gonodactylus espinosus, known only from a single specimen of small 
size, differs from spinosus solely in the greater breadth of the telson and in the total 
absence of spinules on its dorsal surface. In G. demani the variation in the number of 
spinules, as illustrated in the material examined, is so great that should a wholly smooth 
example be added thereto the series would still be almost continuous in its variation. 
On analogy therefore with its ally it may be expected thata complete transition between 
spinosus and espinosus will ultimately be found and, in giving the latter name admission 
as a variety, it must be understood that with the access of fresh material there is a pro- 
bability that the name will cease to possess any greater significance than ‘ incipiens,’ 
‘ tumidus ’ and others which have been employed in the case of G. chiragra. 
In his inclusion of spinosus and espinosus as varieties of G. chiragra Lanchester 
has not, I believe, been followed by any subsequent writer, and I am unable to find any 
sufficient reasons for such a course either in his work or from an examination of the 
material in the Indian Museum. On the contrary the separation of G. chiragra and its 
variety from G. demani and its variety has been conspicuously easy in performance. 
Gonodactylus festae, Nobili, is an Atlantic species resembling G. oerstedi in the 
possession of a short additional carina on the intermediate marginal spines of the telson, 
but differing in the spinulation of this segment. It is only known from the original 
description based on a small number of specimens. It bears, perhaps, much the same 
relation to G. oerstedi that G. demani does to G. chiragra, though, according to Nobili’s 
account, it differs from all these and resembles de Man’s acutirostris in the sharply pro- 
duced antero-lateral angles of the rostrum. 
Gonodactylus glabrous, Brooks, and G. graphurus, Miers, are more clearly dis- 
tinct and in the possession of five long keels in the middle of the dorsal surface of the 
! See addendum, p. 198. 
