_ 
15 
which description of evidence has, I am happy to say, been more 
fully developed and firmly established by the talented coadjutor of 
Prof. Owen, Mr. Quekett of the Royal College of Surgeons, who has 
publicly taught it in the Theatre of that Institution without question 
or contradiction of its truth. This great radius and ulna in Mrs. 
Smith’s Collection I referred to my previously established species, P. 
giganteus, believing at that time that they were probably the bones 
of a fully developed animal, while those previously described were 
the remains of animals not developed to the full extent of their capa- 
bility. 
Sines the publication of these specimens it has been my good for- 
tune to obtain the snout of another and still larger species of Pte- 
rodactyl, from the same pit at Burham in Kent, and which it is 
probable will ultimately prove to belong to the species to which the 
enormous pair of bones in the Cabinet of Mr. Charles of Maidstone 
belongs. Should this hereafter prove to be the case, it will then re- 
main to be shown whether the beautiful specimen of radius and ulna 
in the Collection of Mrs. Smith of Tunbridge Wells, and the bone 
nearly corresponding in size with them, and which was in the possession 
of the Earl of Enniskillen, belong to the newly discovered species, 
which I purpose designating Pterodactylus Cuvieri, or to the pre- 
viously named species, P. giganteus ; or whether there be yet a third 
species existing in the chalk, to which these bones of an intermediate 
size may hereafter be referred *. 
The snout of the new species, P. Cuvieri, differs materially in its 
form from the same part of P. giganteus: while the latter agrees as 
nearly as possible in that respect with P. crassirostris and P. brevi- 
rostris, the former appears to approach very closely the proportions 
of P. longirostris. Thus, if we take the length of the snout from the 
distal end of the cavitas narium, as compared with its height, at the 
same point of P. crassirostris, P. brevirostris and P. giganteus, we 
find the relative proportions to be,—of the first-named, 29 of height 
to 56 of length; of the second, 28 of height to 50 of length; and of 
the third, 28 of height to 58 of length ; we may therefore reasonably 
conclude that, when perfect, the head of P. giganteus very closely re- 
sembled in its proportions that of crassirostris. The length of the 
fragment of the snout of P. Cuvieri at the upper portion of the head 
is 7°20 inches ; at the palatal bones, 6°38 inches; and in this space 
there are sockets for twelve teeth on each side. The distance between 
each tooth is about 14 of the long diameter of the sockets, which are 
somewhat irregularly placed, but are nearly equidistant from each 
other. The pair of teeth at the distal end of the snout appear, both 
from the position of the sockets and the tooth remaining in situ, to 
have been projected more or less forward, in a line with the palatal 
bones. The head appears to have been exceedingly narrow through- 
out the whole of its length. At the third pair of teeth from the distal 
* A third species, C. compressirostris, has since been described by Prof. Owen, 
page 95, Part III. of ‘The Fossil Reptilia of the Cretaceous Formations,’ pub- 
lished by the Palzontographical Society, and to which species the bones in ques- 
tion have been referred. 
