Dorsey: VARIATION IN FLORAL STRUCTURES OF ViTIs 49 
forms are shown in PLATE 2, FIG. 15-18; these show different 
stages of stigmatic development occurring on different vines. The 
seed coats in the ovules show considerable development in PLATE 
2, FIG. 14-18. The material, with the exception of PLATE 3, 
FIG. 21-24, was all fixed just after the flower was completely open, 
so there is probably not more than one or two days’ difference in 
their relative ages; the others were fixed before blooming, to show 
the relative position of the anthers, filaments, and pistils in the 
bud. The crouched position of the filaments will be noted jn 
PLATE 3, FIG. 22. In many flowers, however, the filaments are 
€ven more crouched and bent than in this instance. 
SOME ABNORMALITIES OF THE PISTIL: Cases are rarely observed 
in which one or more anthers of a flower may be found with the 
filament adhering to the side of the pistil, the tissue of both 
filament and pistil being united. In this way the stigmatic tissue 
and the anthers, sometimes partly abortive or deformed, are 
brought into very close contact. Some abnormalities of this nature 
have been observed where the stigmatic and anther tissues are 
intermingled, the pollen being to all appearances normal. This 
adherence is probably due to a lack of differentiation in the 
meristem and does not seem to be a case similar to that observed 
by Chamberlain (’97) in Salix petiolaris, in which microsporangia 
were found in the placenta of the ovary, as well as stigmas de- 
veloping on stamens. 
Some vines bear flowers with distinctly pinkish stigmas. The 
writer has observed this in a few wild vines of both V. vulpina 
and V. bicolor, and also in a number of cultivated varieties, as 
in the R. W. Munson. The stigma is distinctly lobed in some 
Vines, both cultivated and wild, the lobes generally corresponding 
to the carpels. In varieties like the Goff, where many grapes on a 
cluster show distinct sutures, varying in number from one to three 
or four, this tendency can be seen soon after blooming; and in a 
few extreme cases the pistil may be nearly separated into two 
Parts. In outline the stigmatic surface varies much, being oval, 
flaring or flat. Under favorable conditions the stigma may become 
receptive before the corolla opens. This condition was observed 
by the writer in both Concord and Hubbard Seedless. 
The writer wishes to acknowledge his indebtedness to Dr. 
"Ne, Sol. Baracr 
4913 
