88 BRAINERD: HYBRIDS OF THE PALMATA GROUP 
Spring Valley, N. Y. One showed plainly a glabrous specimen 
of V. palmata in flower. By the kindness of Miss Kittredge the 
live plant was presented to me in June. On collecting close- 
fertilized seeds the following August, I found on the average 16 
seeds to the capsule; that is, about three fourths of the ovules 
were undeveloped. Among the offspring raised the following 
season, were plants representing not only the deeply lobed leaves 
of V. palmata but also the uncut leaves of V. papilionacea. Both 
forms, however, were glabrous, as was to be expected; for a 
dominant character like pubescence, when once lost, never reap- 
pears. Nevertheless, the hybrid origin of the wild plant was 
clearly established. Let me cite one further instance. 
On April 15, 1909, I found at Tryon, N. C., what seemed to be 
a colony of this glabrous V. palmata. In the vicinity grew both 
the normal hairy V. palmata and the glabrous V. papilionacea. 
Of the anomalous plants six with leaves moderately lobed were 
shipped to Vermont to be grown and studied in the garden. All 
proved to be more or less infertile, averaging about 20 seeds to 
the capsule. Offspring were raised from all six; and in each brood 
but one were to be seen plants with uncut leaves, indicating that 
one parent of the original hybrid was V. papilionacea; from which, 
doubtless, was also inherited the glabrous character of the plants 
at Tryon. 
Viola palmata X triloba hyb. nov. 
First 1-3 leaves of spring often uncut as in V. triloba, followed 
by others more or less 5—7-lobed, much like those of V. palmata 
but the basal lobes broadly dilated and coarsely toothed as in 
V. triloba; later leaves less dissected, mostly 3-lobed or obscurely 
lobed; capsules infertile, about one third of the ovules maturing 
into seeds; offspring multiform, often bearing on the middle 
segment of the trilobed leaves 4-8 undulations or coarse teeth. 
The genuine V. Angellae Pollard (Torreya 2: 24. 1902), 
proves to be this hybrid, as the original description and the type 
sheet of mature plants (U. S. National Herbarium no. 352,093) 
clearly show. One of the flowering plants on sheet no. 364,862 
may, however, be pure V. palmata. It would seem that the 
original collection by Miss Angell, June 1899, contained many 
specimens of V. palmata, as some of them are to be seen mounted 
