BRAINERD: HYBRIDS OF THE PALMATA GROUP 89 
in the National Herbarium, and others were distributed only 
last summer with the name “ V. Angellae” on the printed labels. 
Possibly some of these plants may be imperfect reversions to 
V. palmata. F urthermore, as already stated, most of the speci- 
mens of “ V. Angellae”’ exhibited at Bronx Park May 1905 were 
V. palmata. Others also have erroneously assumed that plants 
that. they collected at the type station were therefore typical 
V. Angellae. Mr. Pollard distinctly states that his new species 
is associated with V. palmata in the Orange Mountains. 
In May 1908 Dr. E. L. Greene kindly sent me a dozen plants 
of his V. variabilis from the type station, opposite Harpers Ferry 
and named by him Maryland Heights. Most of these plants have 
since been growing in the garden, and from some of them have 
been raised two generations of offspring. These plants I make 
out to consist of two different hybrids of V. ¢riloba, one being the 
V. palmata X triloba above described. A close comparison of the 
characteristic leaf of this hybrid (PLATE 7, FIG. 3 and 4) with the 
leaves of the supposed parents (FIG. 1 and 2) will disclose its com- 
Promise outline. The leaves of Dr. Greene’s plant, though at 
first Suggesting V. palmata, differ in having the broad basal lobes 
of V. triloba even as early as May 31; while specimens collected 
July 22 and September 16 display leaves decidedly trilobed, and 
some hardly lobed at all. This tendency towards uncut leaves 
i autumn is characteristic of V. triloba but not of V. palmata. 
Furthermore, in some of the segregating offspring (PLATE 7, 
FIG. 5 and 7) may we not see in the oddly shaped middle segment 
the resultant of opposite impulses, one demanding 9-11 lobes, the 
other forbidding more than 3? 
So much for the evidence from the living plant, cultivated for 
r s€asons and through three generations. The conclusion 
drawn from this experiment is well sustained by a careful study 
of the seven sheets of “ V. variabilis” at the National Museum. 
. Four were collected May 14, 1898—two by Dr. Greene, and two 
by Mr. Pollard, who accompanied’ him—and three others by 
tT. Greene May 10, 1903; all from Maryland Heights oppo- 
Site Harpers Ferry. The 17 plants here mounted I regard as a 
medley of at least five distinct forms, three of which are repre- 
_ Sented in the live plants sent in 1908. I shall here speak of only 
fou 
