276 BROADHURST: STRUTHIOPTERIS IN NORTH AMERICA 
the larger, more open laminae, 1-5 cm. long, 4-10 mm. wide, often 
opposite, especially near the base; margins entire, irregularly 
revolute in some of the smaller, heavier fronds only; leaf tissue 
membranous to coriaceous as previously described, a few of the 
thinner ones somewhat punctate as in S. exaltata, without scales, 
except rarely a few minute fibrillose ones on the rachis; veins* 
neither raised nor grooved, the apices but slightly or not at all 
glandular, vein spaces 10-15 to I cm. Sporophyls 38-120 cm. 
long; stipes 11-60 cm. long, often marked for a short distance 
by vestigial pinnae, darker than the sterile; lamina 19-60 cm. 
long, 4-10 cm. wide, very gradually reduced at the base and at 
the apex; pinnae 40-50-jugate, 2.2-5 cm. long, 2-3 mm. ide, 
the apex acute, with or without a contracted sterile tip I-2 mm. 
long, the base decidedly dilated; indusium membranous, usually 
entire or subentire, sometimes irregularly broken, never becom- 
ing lacerate; sporangia dark brown. 
DIstTRIBUTION: Pacific coast from California to Alaska. 
SPECIMENS } INCLUDED: CALIFORNIA: Humboldt County, damp 
shady woods, Kellogg & Hartford 1175. OREGON: Nehalum, creek 
banks and dry woods, Kirkwood 110. WASHINGTON: Near 
Tacoma, shady ravines and springs, Flett 2030. BrrtisH COLUM 
BIA: Revelstoke, woods, altitude 1,600 ft., Shaw 835. Stevens Pass, 
altitude 1,150 m., Sandberg & Leiberg 774. ALASKA: Windham 
Bay, Culbertson 4936. WKetchikan, near stream leading from 
Ketchikan Falls, Broadhurst ror. 
Two fronds collected by Douglas in the interior of northwesterm 
America (exact locality not given) formed the type of Hooker s 
Blechnum doodioides; the half sterile, half fertile leaves, with their 
occasionally forked pinnae indicate an abnormal condition, and 
could not be considered as characterizing a valid species. Sim 
ilar abnormal forms are not uncommon in other species of 
genus. The veining of the fertile pinnae and the short, often 
unconnected sori suggest Woodwardia or Doodia. The costal a 
are characteristic of Blechnum, but there are no Blechnums in this 
region. Abnormal fronds in other species often show great YEE 
*The veins are normally once forked; but the crenata variety is often ge 
forked; if the margin is deeply crenate, one main vein may have two OF three 
i Another variation in the veining is found in Umbach 611 from es in 
mish, Washington, where an apparently normal S. Spicant has anastomosing ee 
the lower half of many of the pinnae; the veins form one row of areolae aloné 
midrib, and do not branch again before reaching the margin. 7) Coe 
+ These specimens are all in the herbarium of the New York Botanical © 
