Davis: LAMARCK’S EVENING PRIMROSE 525 
istic of the younger flowering shoots of De Vries’s Lamarckiana. 
On the contrary the bracts are narrow and short-petioled and the 
inflorescence is more open, in agreement with grandiflora. 
3. Bups. The buds are not stout as in the Lamarckiana of 
De Vries and the sepal tips are much more attenuate, a distinctive 
character of grandiflora. The form of the buds is exactly as in 
grandiflora, and this character is one of the most important points 
of agreement with this species. The pubescence on the sepals, as 
described by M. Gagnepain, is short; there are not present the 
numerous long hairs from papillae, which are characteristic of De 
Vries’s Lamarckiana. : 
4. FLowers. The flowers have the very long and delicate 
hypanthium characteristic of grandiflora. Miss Eastwood reports 
that the petals in a bud dissected by her are entire. This is a 
character typical of neither grandiflora nor the Lamarckiana of De 
Vries, but as stated before, the petals of these forms vary so greatly 
in the degree of their indentation that the character has little if any 
taxonomic value. The length of the petals, between 3 and 4 cm.., 
is that of grandiflora and also of certain forms of De Vries’s 
Lamarckiana. The style extends beyond the tips of the anthers 
so that the lobes of the stigma (s, in PLATE 37) are above the latter 
and could not be pollinated in the bud. In these respects the 
flower agrees with both grandiflora and the large-flowered types of 
Lamarckiana in De Vries’s cultures. 
5. CapsuLEs. There are apparently no mature capsules on 
the specimen, so direct comparisons are impossible. Since the 
size and form of a capsule depends upon the development of the 
ovules, i. e. upon whether or not the stigma has been fully polli- 
nated, it is unsafe to accept statements of size unless there is 
evidence that sufficient material has been examined. Lamarck’s 
statement that the capsules are short was probably based on 
immature or partially pollinated capsules. His description of the 
capsules as glabrous points to grandiflora; as stated before, the 
capsules of De Vries’s Lamarckiana are decidedly puberulent and 
Pilose. 
The characters of Oenothera grandiflora, which appear on the 
herbarium sheet of Lamarck’s plant, and those of the Lamarckiana 
of De Vries’s cultures may be more readily contrasted in the 
following statement. 
