194 



D. H. Scott. 



of the glands that my friend Prof. F. W. Oliver first identified ^the 

 seed of Ltjf/inodendron.^) 



The seed named by Williamson (in MS.) Lagenostoma Lomaxi 

 (Fig. 23j is enclosed in an outer envelope or cupule, which bears 



Fig-. 23. Lagenostoma Lomaxi. the seed of Lyginodendron oldhamium, in lougi- 

 tudinal section, enclosed in the loose cupule, bearing capitate glands. X about 15. 



numerous capitate glands identical in structure and form with those 

 on the vegetative organs of Lißjinoäendron oldhamium, with which 

 it is constantly associated (compare Figs. 25 & 26). No other fossil 

 plant is known with any similar appendages, and the evidence of 

 the glands alone would be amply sufficient to justify the attribu- 

 tion, considering the close association of the vegetative and repro- 

 ductive organs in question. Further anatomical evidence is supplied 

 by their internal anatomy. The vascular bundle of the pedicel bearing 

 the seed has the same structure as that of a small rachis of Lygino- 

 dendron. while the smaller bundles which traverse the cupule agree 

 with those in the lamina of the vegetativ^e leaflets. 



The cupule of the seed of Lyginodendron was a deeply-lobed en- 

 velope, which we have compared to the husk of a hazel-nut (see Figs. 

 23 & 24). It overtopped the seed, and enclosed it in the young condi- 

 tion. The pedicel bearing the seed is traversed by a concentric vas- 

 cular bundle, which, before entering the chalaza, gives ofi* numerous 

 branches into the cupule. 



^) Oliver and Scott, On Lagenostoma Lomaxi, the Seed of Lyginodendron, 

 Proc. Eoyal Soc, Vol. 71, 1903. On the Structure of the Palaeozoic Seed Lagenostoma 

 Lomaxi, Phil. Trans. Royal Soc. (B), Vol. 197, 1904. 



