RELATIONSHIPS OF THE AUSTRALIAN CAINOZOIC SYSTEM. 
that Douvillé has pointed out* that the Lepzdocyclinae fall into two 
groups :—Ist. The L. dilatata group, in which the vertical pillars are 
small and uniformly distributed over the shell, the test being 
typically large, as L. chaperi, L. insulae-natalis, L. verbeeki, and L. 
elephantina. 2nd. The L. marginata group, in which the pillars are 
more or less developed, but always more crowded towards the centre 
of the test, as L. raulini, L. morgani, L. tournoueri, L. submarginata, 
and L. sumatrensis. 
The beds in Borneo, Italy and Panama (San Juan), characterized 
by the first group, that of L. dilatata, belong to the Aquitanian stage. 
The beds in Borneo, the south of France and Panama containing 
those of the second, L. marginata group, belong to the Burdigalian 
stage. 
The distribution of the Victorian forms of Lepidocyclina may 
best be shown by the following schedule :— 
Victorian Localities. Species. Beds elsewhere. 
Batesford .. .. |L. tournouert, L. marginata,| Burdigalian of southern 
L. martini Europe 
Keilor be .. |Z. tournouert, L. verbeeki ..| Gaj Beds of India; Upper 
Aquitanian, S. of Europe ; 
L. insulae-natalis Beds of 
Christmas Id. 
Clifton Bank, Muddy | L. verbeeki me .. | Lower Aquitanian 
Creek 
F. Sacco has studied the faunas containing Lepidocyclina and 
Miogypsina with especial regard to the Tertiary basin of Piedmont, 
Italy ; and, although he differs from Douvillé and Prever with 
reference to the precise horizon of L. marginata in that.area, yet that 
question does not affect our present conclusions. M. Sacco fixes 
the L. marginata beds as Aquitanian (but Miocene), whilst Douvillé 
and Prever place them in the Burdigalian (still Miocene). The 
succession remains the same, and the periods follow suit. It is thus 
merely a local adjustment of terms.t 
Prof. A. Silvestri, in his “ Distribuzione Geografica e Geologica 
due Lepidocicline communi nel Terziario Italiano,’§ cites 
the occurrence of both ZL. dilatata and L. tournouert in the 
Priabonian (Oligocene) in Italy and Greece, and their recurrence 
in Italy in the Miocene. In the former instance those species are 
associated with more archaic forms, as the striated nummulites 
and Chapmania, which, however, are absent from the Australian 
Tertiaries. 
* Bull. Soc. Géol. France, sér. 4, vol. vii., 1907, p. 57. 
+ H. Douvillé. Bull. Soc. Géol. France, sér. 4, vol. v., 1905, p. 454 (Table), and p. 455. 
t “Sur la Valeur Stratigraphique des Lepidocyclina et des Mioeypena.? Bull. Soc. Géol. 
France, sér. 4, vol. v., 1906, p- 882. 
§ Mem. Pont. iNoenel Rom! dei Nuovi Lincei, vol. xxix., 1911, pp. 54, 55. 
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