The Non-Marine Mollusca of Portuguese East Africa. 135 



The slender vas deferens runs forward beside the female duct almost 

 to the genital atrium ; it then bends round towards the penis and immedi- 

 ately enlarges to form the somewhat broader and extremely long epiphallus. 

 At the junction of the vas deferens and the epiphallus the latter organ 

 bears a minute oval calciferous sac or flagellum, so small that it might 

 easily escape notice if it were not for its opaque white colour. The epi- 

 phallus passes backwards for nearly 6 mm. and then bends forwards again, 

 giving off at the angle a rather long slender caecum. The penial retractor 

 unites with the epiphallus just in front of the origin of this caecum, which 

 seems also to be attached to the retractor at two points near its anterior 

 end. The epiphallus then passes forwards to unite with the posterior end 

 of the penis, which is nearly 4 mm. long and somewhat swollen, but has a 

 slight constriction in the middle. Its walls are longitudinally folded 

 within, and lined with very minute papillae. Towards its posterior end the 

 penis bears on one side an oval glandular appendage, enclosed in a very 

 thin sheath which surrounds also the extremity of the penis and the anterior 

 end of the epiphallus. (This sheath is not shown in the figure.) 



Semper and Godwin-Austen both figure a somewhat similar penis, but 

 the German author states, possibly in error, that there is a " caecum musculi 

 retractoris penis," in addition to the glandular appendage, the long caecum, 

 and the very small calciferous sac. Godwin- Austen's drawings depict a far 

 larger calciferous sac than that found in the animals from the Lebombo Moun- 

 tains. Pilsbry's figures of the reproductive system of Ledoulxia mesogaea 

 (Mts.) and L. lessensis Pilsbry * show that these species have the same 

 type of genital organs as that described above, although they differ in detail. 



The shells from the Mtisherra R. Valley differ rather noticeably from 

 type. The whorls are tighter coiled, there being about three-quarters of a 

 whorl more in a diameter of 12 mm. than in mozamhicensis , and the sides 

 of the spire are a little concave near the apex, so that the 2 apical whorls 

 are somewhat mammillate, whereas in mozambicensis the sides are convex ; 

 the transverse striation also appears very slightly stronger than that of 

 mozambicensis. 



Ledoulxia albopicta (Mts.), 1869. 



1869. Nanina mossambicensis Pfr., var. albopicta Mts., von der Decken's 

 Reisen, iii, p. 56, pi. 1, fig. 2. D.F. 



1885. Trochonanina anceyi Bgt., Helixarionidees, p. 9. D. 



1897. ,, jenynsi Pfr., var. subjenynsi Ancey, Mts., D.-O.-A., 



iv, p. 49. D. 



Hab. Mozambique (in coll. Ancey). 



L. Marques. Tette (Peters). 



* Bull. Amer. Mus. Nat. Hist., vol. xl, 1919, p. 246, fig. Ill, p. 249, fig. 114. 



