106 THE VOYAGE OF H.M.S. CHALLENGER. 
true glandular part and of a wider part, which acts probably as a receptaculum and opens 
into the fine pore." 
3. The glands in the fourth joint of the legs of the males. These I observed in the 
following species :—Nymphon hamatum, Nymphon brachyrhynchus, Ascorhynchus glaber, 
Colossendeis leptorhynchus, Colossendeis proboscidea, Colossendeis megalonyx, Oorhynchus 
aucklandie, Phoxichilidium insigne, and Pallene australiensis. Whether or not they 
occur in the thighs of the males of all the species I dare not assert. I can only say that 
I did not find them in the thighs of the following species:—Nymphon robustum, 
N. brevicaudatum, N. longicoxa, N. grossipes, and Colossendeis gracilis. This may, 
however, be the consequence of these glandular masses being present only during a short 
period of the year—for example, only in the breeding season. 
These are the glands which were observed by Dohrn. According to him (loc. cit., 
p- 36) they occur only in the male sex, and are found in the fourth joint of the legs, viz., 
in those joints in which in the females the ovaries are most strongly developed. Dohrn, 
moreover, tells us that these glands are extremely variable in their appearance ; while in 
Ammothea they are furnished with a single duct only, in Phowichilus there are fifteen 
smaller openings. Dohrn supposes that the function of these glands is to secrete a viscous 
fluid, wherewith the males agglutinate the eggs which have been laid by the females, 
and attach them to their ovigerous legs. 
As to the function of these glands, I was unable to make any observations from the 
material preserved in spirits and brought home by H.M.S. Challenger. With regard to 
their structure the following may be stated:—The glands are always composed of a 
skeleton of connective tissue, the meshes of which are or are not filled with nucleated 
cells, which doubtless are the true glandular cells. When these glands have the meshes 
filled up with the nucleated cells (Pl. XVI. figs. 5 and 10), the skeleton of con- 
nective tissue is not easily discerned; while in those cases in which the meshes 
are empty (Pl. XVI. fig. 15) the structure of the connective tissue is easily observed. 
In the form of the glandular cells small differences were also observed. In Ascorhynchus 
glaber, they are fusiform and pointed at both extremities; in Mymphon hamatum, 
they are rounded, but in both cases distinct nuclei are present. A considerable 
difference, moreover, is seen in the greater or less degree of concentration which 
the glandular masses had undergone. In Nymphon hamatum and in Ascorhynchus 
glaber the gland in the fourth joint (which, however, in the latter species is also 
present in the fifth joint of the leg) forms only a single mass, which runs through 
the whole joint and opens at the one side in a row of pores, each of which is placed at 
the tip of a chimney-like process. As seen in fig. 6, the gland in Nymphon hamatum 
almost extends on the one side of the leg, between the intestinal caecum and the 
1 These glands are also observed by Dohrn ; he calls them “ ein zweites noch grésseres Excretions- (oder Driisen ?-) 
Organ (Ibid.). 
