PROGRESSIVE METAMORPHOSES. 291 



pistil, and not adherent as in the case of Musscenda. In both 

 instances it will presumably be purely axial in character. 



Progressive changes in the calyx are not uncommon by 

 its assuming a petaloid character. This is normal in some 

 genera of Banunculacece, in Fuchsia, BhodocJiiton, as well as in 

 some members of the Incompleted, as in Mirahilis, Folygonuw, 

 Daphne, etc. ]S"ormally coloured sepals are most frequent 

 in polysepalous genera. Abnormal colorisation, with or 

 without any metamorphosis of the organ, is 

 most frequent in gamosepalous flowers, as in 

 the cultivated " hose-in-hose " varieties of 

 Primula, Miynidus and Azalea. The calyx 

 may be petaloid either wholly or in part 

 only. In Musscenda (Fig. 68), one sepal only 

 is normally sub-petaloid. Calceolaria has 

 occasionally one or more sepals petaloid. 

 Similarly Linaria (Fig. 69) and other in- 

 stances might be mentioned. These condi- 

 tions, brought about by cultivation, clearly Fig. G^unaria. 

 show the important part that high nourish- petaloid. 

 ment plays as an external stimulus or factor in the produc- 

 tion of colour. 



Staminoid sepals appear to be very rare. It is recorded 

 by M. Gris that they have occurred in Philadelphus speciosus:* 



Pistiloid sepals are nearly equally as rare as staminoid. 

 They have been observed by Mr. Laxton in double flowers 

 of the Garden Pea (Fig. 70), in which there was a five or 

 six-leaved calyx, some of the segments of which were of a 

 carpellary nature, and bore imperfect ovules on their mar- 

 gins, the extremities being drawn out into sub-stigmatiferous 



styles.t 



* Bull. Soc. de Bot. Fr., t. v., p. 330. 



t Gard. Chron. 1886, p. 897 ; and Teratology, p. 302. 



