228 Harris: Anomalous anther-structure 



gonum concretis intus 5 -locularibus, loculis nempe 3 externis, 2 

 intern is regulariter dispositis ! M but Grisebach writes: "The 

 anomalous structure of the column, described by DeCandolle, 

 must have been a monstrosity; for I find 10 anther-cells, regularly 

 arranged in a single row." The figure given by Schnizlein rep- 

 resents a condition such as DeCandolle describes. Schumann 

 says of the androecium : " Stb. unter sich der ganze Lange nach 

 zu einer dem Grunde der Blkr. angehefteten Rohre verbunden ; 

 A. nur in der oberen Halfte, wie mit Poren geoffnet." 



Serial sections represented by figure 3 show the real condition 

 of this synandrium. At the tips only the outer wall maybe seen, 

 smooth externally and rough internally. As the sections are fol- 

 lowed down towards the base the cone is seen to be formed by the 

 fusion of five 4-locellate anthers. Between these there are towards 

 the outside four intercellular spaces "which have been mistaken for 

 locelli, and so lead to the statement that the synandrium has in 

 the outer row fifteen and in the inner row ten locelli. 



Missouri Botanical Garden. 



