Hepaticae of Puerto Rico 
IV. ODONTOLEJEUNEA, CYCLOLEJEUNEA AND PRIONOLEJEUNEA 
ALEXANDER W. EvANs 
(WirH PLaTEs 8-12) _ 
ODONTOLEJEUNEA 
In his subgenus Odonto-Leyeunea, Spruce * grouped together 
a number of Lejewncae Holostipae in which dentate or spinose 
leaves were borne on a prostrate axis, the latter being closely ad- 
herent to the substratum by means of radicelliferous discs devel- 
dope on the underleaves. In the other subgenera of this group, 
in which dentate or spinose leaves also occurred, as, for example, 
Thysano-Lejeunea and Dicrano-Lejeunca, the leafy axes were 
pendulous or ascending from a prostrate caudex and were almost 
or quite destitute of rhizoids. Odonto-Lejeunea was further char- 
acterized by a strongly flattened perianth more or less toothed 
along the sharp lateral keels and emarginate at the apex. Although 
classed among the Holostipae, Spruce included in his subgenus 
certain species, such as Lejeunea Peruviana Lehm. & Lindenb., 
in which the majority of the underleaves were bifid, and others, 
such as L. convexisti~a Lehm. & Lindenb., in which a tendency 
toward the development of bifid underleaves might sometimes be 
observed. Schiffner’s genus Odontolejeunea + has the same limits 
as Spruce’s subgenus,but Stephani understands the genus ina some 
what more extended sense and includes in it at least one species, 
L. accedens Gottsche, t which Spruce referred to Priono-Lejeunea 
(under the name Z. leptocardia Spruce). In this species, as well 
as in certain others recently described by Stephani,§ the under- 
leaves are invariably bifid. Although in many respects a natural 
group, the genus Odontolejeunca as understood by Stephani or 
even by Schiffner is more comprehensive than is usual among the 
oo 
* Hep. Amaz. et And. 142. 1884. 
t Engler & Prantl, Nat. Pflanzenfam. 13: 127. 1893. 
{ Hedwigia, 27: 284. 1888 
4L. c.35: 120. 1896. 
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