THE CHORDA TYMPANI. 171 



rest of the nerve) clearly is a visceral nerve for the 

 mucosa of the month, the r, mandibularis internus VII of 

 Stannius in the selachians and of Strong- for the Amphibia. 

 In Stannius' discussion of the homologue in the selachians 

 of the r. hyoideus of teleosts (p. 65) I find no statement 

 which would identify the "internal mandibular" with a 

 motor nerve in either selachians or teleosts, and there is 

 no reason why that term should not be applied to the 

 communis component of the r. mandibularis VII of the 

 bony fishes as in other vertebrates. 



Cole follows (p. 202) with a tabular presentation of his 

 conception of the composition of the facial nerve, a con- 

 ception which I think inadmissible in several particulars. 

 My own view of the branchiomeric characters of the 

 facial nerve is given under the caption Metamerism in 

 Section 12. 



As these sheets pass through the press (July, 1899), I am con- 

 strained to add a further note. Cole has admitted the misquotation 

 of Stannius, both to me privately, and later publicly in the Anato- 

 mise her Anze/^-er (XVI, 2, 1899, p. 40, ff.) Since the matter is of 

 some morphological importance, I have, however, left the criticism 

 as originally written, in order that Stannius' own words might be 

 before us. 



Another matter may be touched upon here. In the course of a 

 microscopical review of the trigemino-facial complex of Gadus 

 morrhua, upon which I am now engaged, I find that the post- 

 spiracular communis element of the facialis is totally wanting (or at 

 most, so reduced as to be unrecognizable), z. <?., there is no r. mandib- 

 ularis internus VII in Gadus (Cole's statement to the contrary, '98a, 

 p. 202, notwithstanding). The nerve which I have called the r. pre- 

 trematicus VII is present in exactly the same relations as in Menidia, 

 and there is t'n addition a large communis element in both the r. 

 maxillaris and the r. mandibularis V. The mucosa in the region of 

 the suspensorium and of the mandible which is supplied from the r. 

 mandibularis internus VII in Menidia is supplied from the mandib- 

 ularis trigemini in Gadus. 



In the light of the preceding discussion it would therefore appear 

 probable that in Gadus the r. pre-trematicus VII represents the 

 nerve i of Stannius in selachians and that the nerve 2 (/. e., the 

 chorda tympani) has fused with the r. mandibularis V. In any case 

 this emphasizes the difficulty alluded to above of defining the homol- 

 ogies of these nerves by topographical relations alone. It also makes 



