APPENDIX—HELICIGONA ARBUSTORUM., A491 
Variation.—Dr. P. Germain, who has especially studied the effect of 
environment upon shell form, affirms that //. arbustorum and other species 
when living under exceptionally humid conditions become remarkable for 
the slow coiling of their shells, and by the compression of the last whorl, 
and infers from the presence of these characters in the fossil specimens of 
H. arbustorum from the various Pleistocene deposits of the south of France 
that the locality they inhabited was of a very humid character. 
Dr. Germain has described and figured (Archiv. Nat. Hist. Mus. Lyons, 
1912, pp. 72 and 166-170, pl. iv., f. 148) a new variety of this species, 
which is said to be remarkable for its very depressed form, thick, solid, and 
somewhat cretaceous shell, which is finely striate as in var. minor Locard, 
and has a very narrow supra-carinal brown band. 
He also notes from the deposits about Lyons three distinct forms of 
this species, which he differentiates as major Locard, intermedia Germain, 
and minor Locard. 
The sub-var. minor Locard has a diameter of 14-16 mill., and an altitude of 
11-12 mill., and was common about Lyons at the time of the deposition of the 
loess, becoming successively modified into the vars. intermedia and major as the 
conditions became less rigorous, the var. major being now the typical form of the 
present day about Ly ons. 
The sub-var. intermedia Germain is 19-20 mill. diameter, and 15-16 mill. in 
altitude, thick, quite solid, and somewhat heavy, with a globosely-elevated form. 
Dr. Gaillard found specimens in the loess at L’Ile Barbe, St. Rambert, Rhone, and 
at Béeeude, in the commune of Feyzin, sere. 
The sub-var. major Locard, from the tufa of Buisse, Isere, is 22-234 mill. in 
diameter and 16-173 mill. in altitude,.and is the form now occupying the plains, 
and is rare about Lyons. 
Var. canigonensis Boubée (see p. 440). 
Helix arbustorum var. dorie Paulucci, Faune Mal. Italie, 1878, p. 31. 
The sub-var. dorige is described as possessing a shell of an uniformly dark 
olive-green colour, completely destitute of the characteristic marblings, but some- 
times showing a slightly perceptible supra-peripheral band. 
PEATE. 
Piedmont—Found by the Marquis Doria on Mont Barone near Biella (Paulucci, 
Faune Mal. Italie, 1878, p. 31). 
: Ave 2 
Var. gaillardi Germain. 
Helix arbustorum var. gaillardi Germain, Moll. terr. et fluy. quatern des bassins du Rhone et du 
Rhin. Archiv. Mus. Hist. Nat., 1912, p. 72, pl. iv., ff. 148, 166-170. 
SHELL size of var. minor Loecard, but much more depressed, spire little risen, 
whorls six, convex, regularly increasing in size, and separated by a well-marked 
suture, last whorl noticeably more convex below than above, and deflected at the 
mouth; mouth transversely oval and quite oblique, reflected over the umbilicus, 
-which it almost entirely covers ; peristome thickened and reflected. 
Diam. max 15mill., min. 13mill.; alt. 9mill.; diam. of apert. Smill., alt. 7 mill. 
Discovered in the loess of L’fle Barbe, St. Rambert, near Lyons, by Dr. Cl. 
Gaillard, to whom the form is dedicated. 
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