DARK BRAHiMA AND TOSA FOWL. 51 



Considering the intrinsic difficulties of classification due to the partial 

 blending of characteristics, there is a fairlj' close correspondence between the 

 calculated and the actual. This result proves that there is little if any 

 necessary correlation between the characteristics in question ; they may 

 combine in a chance fashion in the second hybrid generation. 



CONCLUSIONS. 



The inheritance of color in this cross between a white and a game-colored 

 breed is remarkable in that white is not dominant — as is usually the case — • 

 nor recessive ; but inheritance is particulate in the heterozygote, producing 

 barred offspring. Segregation nevertheless occurs in the second hybrid 

 generation, but the extracted whites and game colored birds are, for the 

 most part, no longer as pure in color as their grandparents were. The 

 germ cells are no longer perfectly pure — they have become infected by con- 

 tact with the opposite quality. 



The long-tailed characteristic behaves in inheritance like a unit character — 

 in no wise different from plumage color. One can not help doubting whether 

 it originated by any different method from that in which the diverse colors 

 of poultry have arisen. 



Foot feathering is dominant here as in many other cases ; yet the domi- 

 nance is incomplete. The germ cells of the .second hybrid generation are 

 no longer pure. 



The White Cochin has no sexual dimorphism in plumage color, while the 

 Tosa fowl is strongly dimorphic. Every one of the first hybrids is dimorphic 

 in plumage coloration, the two .sexes resembling, except for the white, 

 respectively the female and the male Tosa fowl It is striking to see how 

 from a germ cell of the male Tosa fowl either a bird colored like a male 

 Tosa or a bird colored like a female Tosa may arise. The male germ cells 

 contain the Anlagen not only of the male characteristic but also of the 

 female characteristic (Darwin, 1876, Chapter XIII). 



Series X. — Dark Brahma and Tosa Fowl. 



STATEMENT OF PROBLEM. 



This series was undertaken primarily to test inheritance of secondary sexual 

 characteristics and the possibility of transferring them from one sex to 

 another. 



THR RACKS AS A WHOLE. 



The Dark Brahma male and female have been described at page 32 ; the 

 Tosa fowl, male and female, at pages 43, 44. Each race has a strongly 

 marked sexual dimorphism in plumage color. The males have feathers of a 

 more uniform color ; the female Dark Brahma has penciled feathers ; the 

 female Tosa fowl has mossy feathers with prominent light shafting. 



